Pleurobranchus

Goodheart et al. (2015)[1] recognized 14 valid species, while Alvim & Pimenta 2016[3] recognized one other species. Pleurobranchus sishaensis is additionally recognized by WoRMS.[4]

Species in the genus Pleurobranchus include:

1. Pleurobranchus albiguttatus (Bergh, 1905)[1][4]
2. Pleurobranchus areolatus Mörch, 1863[1][4] Atlantic sidegill slug
3. Pleurobranchus digueti Rochebrune, 1895[1][4]
4. Pleurobranchus forskalii Rueppell & Leuckart, 1828[1][4] - synonym: Pleurobranchus semperi (Vayssière, 1896)[4] / Oscanius semperi Vayssière, 1896[1]
5. Pleurobranchus grandis W. H. Pease, 1868 [1][5]
6. Pleurobranchus hilli (Hedley, 1894)[1][4]
7. Pleurobranchus iouspi Ev. Marcus, 1984[4][3]
8. Pleurobranchus mamillatus Quoy & Gaimard, 1832[1][4]
9. Pleurobranchus membranaceus Montagu, 1815[1][4]
10. Pleurobranchus nigropunctatus (Bergh, 1907)[1][4]
11. Pleurobranchus peroni Cuvier, 1804[1][4]
12. Pleurobranchus reticulatus Rang, 1832[1][4]
13. Pleurobranchus sishaensis (Zhang & Lin, 1965)[4]
14. Pleurobranchus testudinarius Cantraine, 1835[1][4]
15. Pleurobranchus varians Pease, 1860[1]
16. Pleurobranchus weberi (Bergh, 1905)[1][4]

Goodheart et al. (2015)[1] recognized following synonyms:

• Pleurobranchus lacteus Dall & C.T. Simpson, 1901[4] is a synonym of Berthella lacteus (A. E. Verrill, 1901)[1]
• Placobranchiopsis niveus Verrill, 1901 / Pleurobranchus niveus (A. E. Verrill, 1901)[4] is a synonym of Berthella niveus (A. E. Verrill, 1901)[1]

Nomen nudum (nude name)

• Pleurobranchus gigas Sawaya & Grempel, 1972 ;[6] (nomen nudum[7])

Species brought into synonymy

• Pleurobranchus agassizii MacFarland, 1909: synonym of Berthella agassizii (MacFarland, 1909)
• Pleurobranchus americanus A. E. Verrill, 1885: synonym of Berthella americana (A. E. Verrill, 1885)
• Pleurobranchus atlanticus R.T. Abbott, 1949 : synonym of Pleurobranchus crossei Vayssière, 1897
• Pleurobranchus aurantiacus Risso, 1818: synonym of Berthella aurantiaca (Risso, 1818)
• Pleurobranchus beyrichii (Hilgendorf, 1877): synonym of Mikadotrochus beyrichii (Hilgendorf, 1877)
• Pleurobranchus brevifrons Philippi, 1844: synonym of Berthella plumula (Montagu, 1803)
• Pleurobranchus caledonicus Risbec, 1928 accepted as Berthella caledonica (Risbec, 1928)[4]
• Pleurobranchus californicus Dall, 1900: synonym of Berthella californica (Dall, 1900)
• Pleurobranchus citrinus Rüppell & Leuckart, 1828: synonym of Berthellina citrina (Rüppell & Leuckart, 1828)
• Pleurobranchus contarinii Vérany, 1846: synonym of Pleurobranchus membranaceus (Montagu, 1815)
• Pleurobranchus crossei Vayssière, 1897 accepted as Pleurobranchus areolatus Mörch, 1863 [4]
• Pleurobranchus dautzenbergi Watson, 1897: synonym of Berthella dautzenbergi (Watson, 1897)
• Pleurobranchus dehaanii Cantraine, 1841: synonym of Pleurobranchus membranaceus (Montagu, 1815)
• Pleurobranchus delicatus Pease, 1861 accepted as Berthellina delicata (Pease, 1861)[4]
• Pleurobranchus denotarisi Vérany, 1846: synonym of Pleurobranchus membranaceus (Montagu, 1815)
• Pleurobranchus denotarisi Vérany, 1846: synonym of Pleurobranchus testudinarius Cantraine, 1835
• Pleurobranchus elongatus Cantraine, 1835: synonym of Berthella elongata (Cantraine, 1835)
• Pleurobranchus elongatus Cantraine, 1835: synonym of Berthella aurantiaca (Risso, 1818)
• Pleurobranchus emys Ev. Marcus, 1984 accepted as Pleurobranchus iouspi Ev. Marcus, 1984 [4]
• Pleurobranchus evelinae T. E. Thompson, 1977 accepted as Pleurobranchus areolatus Mörch, 1863 [4]
• Pleurobranchus fleuriausi d'Orbigny, 1837: synonym of Berthella plumula (Montagu, 1803)
• Pleurobranchus forskali: synonym of Pleurobranchus forskalii (Rüppell & Leuckart, 1831)
• Pleurobranchus garciagomezi Cervera, J.L., Cattaneo-Vietti & M. Edmunds, 1996 accepted as Pleurobranchus reticulatus Rang, 1832 [4]
• Pleurobranchus granulatus Krauss, 1848: synonym of Berthellina granulata (Krauss, 1848)
• Pleurobranchus hirasei K. Bába, 1971 accepted as Pleurobranchus peronii Cuvier, 1804[4]
• Pleurobranchus inconspicua (Bergh, 1897): synonym of Pleurobranchaea inconspicua Bergh, 1897
• Pleurobranchus inhacae Macnae, 1962: synonym of Pleurobranchus peronii Cuvier, 1804
• Pleurobranchus inhancae [sic]: synonym of Pleurobranchus inhacae Macnae, 1962: synonym of Pleurobranchus peronii Cuvier, 1804
• Pleurobranchus lesueur de Blainville, 1827: synonym of Pleurobranchus membranaceus (Montagu, 1815)
• Pleurobranchus lowei Watson, 1897: synonym of Berthellina edwardsi (Vayssière, 1896)
• Pleurobranchus luniceps Cuvier, 1817: synonym of Euselenops luniceps (Cuvier, 1817)
• Pleurobranchus mammillatus Schultz in Philippi, 1836: synonym of Pleurobranchus testudinarius Cantraine, 1835
• Pleurobranchus monterosati Vayssière, 1880: synonym of Berthella ocellata (Delle Chiaje, 1830)
• Pleurobranchus ocellatus Delle Chiaje, 1830: synonym of Berthella ocellata (Delle Chiaje, 1830)
• Pleurobranchus patagonicus d'Orbigny, 1835: synonym of Berthella patagonica (d'Orbigny, 1835)
• Pleurobranchus pellucidus Pease, 1860: synonym of Berthella stellata (Risso, 1826)
• Pleurobranchus perforatus Philippi, 1844: synonym of Berthella plumula (Montagu, 1803)
• Pleurobranchus perrieri Vayssière, 1897: synonym of Pleurobranchus forskalii Rüppell & Leuckart, 1828
• Pleurobranchus platei Bergh, 1898: synonym of Berthella platei (Bergh, 1898)
• Pleurobranchus punctatus Quoy & Gaimard, 1832: synonym of Berthellina citrina (Rüppell & Leuckart, 1828)
• Pleurobranchus reesi K. White, 1952 : synonym of Pleurobranchus atlanticus Abbott, 1949
• Pleurobranchus scutatus Martens [in Möbius], 1880: synonym of Berthella martensi (Pilsbry, 1896)
• Pleurobranchus stellatus Risso, 1826: synonym of Berthella stellata (Risso, 1826)
• Pleurobranchus strongi Mcfarland, 1966 : synonym of Berthella strongi (MacFarland, 1966)
• Pleurobranchus tuberculatus Meckel, 1808: synonym of Pleurobranchus membranaceus (Montagu, 1815)
• Pleurobranchus xhosa Macnae, 1962 (possibly a juvenile form of P. peroni)
• Pleurobranchus wirtzi Ortea, Moro & Caballer, 2014 accepted as Pleurobranchus reticulatus Rang, 1832 [4]

Nomen dubium

• Pleurobranchus limacoides Forbes, 1844
• Pleurobranchus verrilli Thiele, 1931
• Pleurobranchus ypsilophora Vayssière, 1898

Taxa inquerenda

• Pleurobranchus disceptus O'Donoghue, 1929
• Pleurobranchus griseus Bergh, 1905
• Pleurobranchus latipes Bergh, 1905
• Pleurobranchus moebii Vayssière, 1898
• Pleurobranchus papillosus (O'Donoghue, 1929)
• Pleurobranchus sculptatus (O'Donoghue, 1929)

• SEM images of the radula can be found at Thompson, T.E.; Bebbington, A. (1973). "Scanning electron microscope studies of gastropod radulae". Malacologia. 14: 147–165.
• Howson, C.M.; Picton, B.E. (Ed.) (1997). The species directory of the marine fauna and flora of the British Isles and surrounding seas. Ulster Museum Publication, 276. The Ulster Museum: Belfast, UK. ISBN 0-948150-06-8. vi, 508 (+ cd-rom) pp.
• Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180–213

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