Squamanita contortipes

Squamanita contortipes is a small mushroom species in the family Squamanitaceae, formerly in the Tricholomataceae. It was originally described in 1957 by American mycologist Alexander H. Smith and Daniel Elliot Stuntz as a member of Cystoderma. Paul Heinemann and David Thoen transferred it to the genus Squamanita in 1973.[1] Discovery of an unusual fruiting of this species where three fruitbodies grew on one, still fertile host pileus which was a species of Galerina proved that Squamanita was a mycoparasitic genus. Photos of this fruiting were published in 1994[2] and immediately republished and highlighted in 1995 in Nature magazine where the original discovery article was featured.[3] The species proved to be the Rosetta Stone[2] for deciphering the parasite from the host in graft-like fruitings. Normally, S. contortipes only forms one fruitbody on each parasitized host and the host normally fails to remain fertile and does not form its own pileus.[1]