Arachniodes

Arachniodes is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I).[1] A number of species in this genus are known as "holly ferns".[2]

Arachniodes
Arachniodes simplicior
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Class: Polypodiopsida
Order: Polypodiales
Suborder: Polypodiineae
Family: Dryopteridaceae
Subfamily: Dryopteridoideae
Genus: Arachniodes
Blume
Synonyms[1]
  • Byrsopteris C.V.Morton
  • Leptorumohra (H.Ito) H.Ito
  • Lithostegia Ching
  • Phanerophlebiopsis Ching

The genus Arachniodes was first published by Carl Ludwig von Blume in 1828, with the single Indonesian species Arachniodes aspidioides.[3] The genus was not widely recognized until Mary Douglas Tindale transferred the two species (Byrsopteris amabilis and Byrsopteris aristata) into it in 1961.[4]

Species

The Pteridophyte Phylogeny Group classification of 2016 (PPG I) considers Arachniodes to have 60 species.[1] As of January 2020, the Checklist of Ferns and Lycophytes of the World accepted 78 species:[5]

  • Arachniodes ailaoshanensis Ching
  • Arachniodes amabilis (Blume) Tindale
  • Arachniodes amoena (Ching) Ching
  • Arachniodes aristata (G.Forst.) Tindale
  • Arachniodes assamica (Kuhn) Ohwi
  • Arachniodes bella (C.Chr.) Ching
  • Arachniodes blinii (H.Lév.) Nakaike
  • Arachniodes carvifolia (Kunze) Ching
  • Arachniodes caudata Ching
  • Arachniodes cavaleriei (Christ) Ohwi
  • Arachniodes chinensis (Rosenst.) Ching
  • Arachniodes coniifolia (T.Moore) Ching
  • Arachniodes cornucervi (D.Don) Fraser-Jenk.
  • Arachniodes daklakensis Li Bing Zhang, N.T.Lu & X.M.Zhou
  • Arachniodes davalliaeformis (Christ) Nakaike
  • Arachniodes denticulata (Sw.) Ching
  • Arachniodes fengii Ching
  • Arachniodes festina (Hance) Ching
  • Arachniodes formosa (Fée) Ching
  • Arachniodes formosissima (Goldm.) Nakaike
  • Arachniodes gigantea Ching
  • Arachniodes globisora (Hayata) Ching
  • Arachniodes grossa (Tardieu & C.Chr.) Ching
  • Arachniodes hainanensis (Ching) Ching
  • Arachniodes haniffii (Holttum) Ching
  • Arachniodes hehaii Li Bing Zhang, N.T.Lu & X.F.Gao
  • Arachniodes hekiana Kurata
  • Arachniodes henryi (Christ) Ching
  • Arachniodes hiugana Sa. Kurata
  • Arachniodes hunanensis Ching
  • Arachniodes insularis W.H.Wagner
  • Arachniodes japonica (Kurata) Nakaike
  • Arachniodes jinpingensis Y.T.Hsieh
  • Arachniodes leucostegioides (C.Chr.) Ching
  • Arachniodes longicaudata Li Bing Zhang, N.T.Lu & Liang Zhang
  • Arachniodes longipinna Ching
  • Arachniodes microlepioides (C.Chr.) comb. ined.
  • Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi
  • Arachniodes miyakei (H.Itô) Shimura ex Nakaike
  • Arachniodes mutica (Franch. & Sav.) Ohwi
  • Arachniodes neopodophylla (Ching) Nakaike
  • Arachniodes nigrospinosa (Ching) Ching
  • Arachniodes nipponica (Rosenst.) Ohwi
  • Arachniodes oosorae H.Itô
  • Arachniodes palmipes (Kunze) Fraser-Jenk.
  • Arachniodes pseudoaristata (Tagawa) Ohwi
  • Arachniodes pseudoassamica Ching
  • Arachniodes pseudorepens Nakaike
  • Arachniodes puncticulata (Alderw.) Ching
  • Arachniodes quadripinnata (Hayata) Seriz.
  • Arachniodes quangnamensis Li Bing Zhang, N.T.Lu & Liang Zhang
  • Arachniodes repens Kurata
  • Arachniodes rhomboidea (Schott) Ching
  • Arachniodes rigidissima (Hook.) Ching
  • Arachniodes sarasiniorum (Christ) Nakaike
  • Arachniodes simplicior (Makino) Ohwi
  • Arachniodes simulans (Ching) Ching
  • Arachniodes sinomiqueliana (Ching) Ohwi
  • Arachniodes sinorhomboidea Ching
  • Arachniodes sledgei Fraser-Jenk.
  • Arachniodes spectabilis (Ching) Ching
  • Arachniodes squamulosa R.C.Moran & B. Øllg.
  • Arachniodes standishii (T.Moore) Ohwi
  • Arachniodes subamabilis Kurata
  • Arachniodes superba Fraser-Jenk.
  • Arachniodes tomitae Sa. Kurata
  • Arachniodes tonkinensis (Ching) Ching
  • Arachniodes tripinnata (Goldm.) Sledge
  • Arachniodes tsiangiana (Ching) Nakaike
  • Arachniodes webbiana (A.Braun) Schelpe
  • Arachniodes wulingshanensis S.F.Wu
  • Arachniodes yakusimensis (H.Itô) Nakaike
  • Arachniodes ziyunshanensis Y.T.Xie

Some hybrids were also accepted:[5]

  • Arachniodes × azuminoensis Fujiw., Y.Matsuda, Yu. Abe & Otsuka
  • Arachniodes × chibaensis Yashiro
  • Arachniodes × clivorum Kurata
  • Arachniodes × ikeminensis Seriz.
  • Arachniodes × kenzo-satakei (Kurata) Kurata
  • Arachniodes × kurosawae Shimura & Kurata
  • Arachniodes × masakii Kurata
  • Arachniodes × minamitanii Sa. Kurata
  • Arachniodes × mirabilis Sa. Kurata
  • Arachniodes × mitsuyoshiana Sa. Kurata
  • Arachniodes × pseudohekiana Kurata
  • Arachniodes × respiciens Sa. Kurata
  • Arachniodes × sahashii Fraser-Jenk. & Kandel
  • Arachniodes × sasamotoi Kurata
  • Arachniodes × takayamensis Seriz.

References
  1. PPG I (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution. 54 (6): 563–603. doi:10.1111/jse.12229.
  2. Arachniodes Flora of North America 26 Jan 2012
  3. Blume, Carl (Karl) Ludwig von; Enumeratio Plantarum Javae fasc. 2: 241. 1828.
  4. Tindale, Mary Douglas; Contributions from the New South Wales National Herbarium 3(1): 90. 1961.
  5. Hassler, Michael & Schmitt, Bernd (January 2020). "Arachniodes". Checklist of Ferns and Lycophytes of the World. 8.20. Retrieved 2020-01-17.
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