Fomes

Fomes is a genus of perennial woody fungi in the family Polyporaceae. Species are typically hoof-shaped (ungulate). New growth each season is added to the margin, resulting in a downward extension of the hymenium.[1] This often results in a zonate appearance of the upper surface, that is, marked by concentric bands of color.

This article is about the fungus. For the inanimate disease vector, see Fomite.

Fomes
Fomes fomentarius
Scientific classification
Kingdom: Fungi
Division: Basidiomycota
Class: Agaricomycetes
Order: Polyporales
Family: Polyporaceae
Genus: Fomes
(Fr.) Fr. (1849)
Type species
Fomes fomentarius
(L.) Fr. (1849)

The name comes from Latin fomes, meaning "tinder",[2] from the use of Fomes fomentarius, also known as the tinder fungus, in making tinder (see amadou).

Taxonomy

Fomes was first introduced by Elias Magnus Fries as a subgenus of Polyporus in his 1836 work Genera Hymenomycetum. He promoted it to generic status in 1849.[3]

Description

Fomes species have perennial, hoof-shaped fruit bodies that attach directly to their substrate without a stipe. The cap surface has a hard smooth crust that ranges in colour from gray to blackish. On the underside of the cap, the pore surface is pale brown with small pores, and brown tube layers. The tough and fibrous context is pale brown. The fruit body becomes stratified after multiple years of growth. Fomes has a trimitic hyphal system, containing generative, skeletal, and binding hyphae. The generative hyphae have clamp connections, while binding and skeletal hyphae have pale brownish pigment that appears when they are mounted in a solution of potassium hydroxide. The cystidioles are fusoid, imbedded, or project somewhat. The basidia (spore-bearing cells) are club-shaped with a basal clamp, and have four sterigmata. The spores are cylindric, large, hyaline and smooth, and are non-reactive to Melzer's reagent.[4]

Species

As of June 2017, Index Fungorum accepts 59 species of Fomes:[5]

  • Fomes abramsianus (Murrill) Murrill (1915)
  • Fomes ajazii S.M.Hussain (1952)
  • Fomes albescens (Lázaro Ibiza) Sacc. & Trotter (1925)
  • Fomes albogriseus Peck (1903)
  • Fomes albotextus Lloyd (1924) – Sumatra
  • Fomes angulus Lloyd (1913)
  • Fomes aratus Sacc. & D.Sacc. (1905)
  • Fomes arctostaphyli Long (1917)
  • Fomes auriscalpioides Henn. (1904)
  • Fomes bomfimensis Henn. (1904)
  • Fomes borealis Lloyd (1915)
  • Fomes borneoensis (Lloyd) S.Ahmad (1956)
  • Fomes bossardii Lucien (1923)
  • Fomes chaquensis Iaconis & J.E.Wright (1953)
  • Fomes clelandii Lloyd (1915)
  • Fomes congoanus Bres. (1913)
  • Fomes crispus Lázaro Ibiza (1917)
  • Fomes extensus (Lév.) Cooke (1885 – Uganda)
  • Fomes fasciatus (Sw.) Cooke (1885)
  • Fomes ferrugineobrunneus Cout. (1925)
  • Fomes fomentarius (L.) Fr. (1849) – widespread
  • Fomes fulvellus (Bres.) Sacc. (1891)
  • Fomes fulvus (Scop.) Gillet (1878)
  • Fomes goethartii Bres. (1910)
  • Fomes griseus Lázaro Ibiza (1916)
  • Fomes haeuslerianus Henn. (1896)
  • Fomes halconensis Bres. (1912)
  • Fomes hemitephrus (Berk.) Cooke (1885) – Australia
Fomes hemitephrus photographed in Tasmania
  • Fomes idahoensis R.W.Br. (1940)
  • Fomes imitator Petch (1922)
  • Fomes javanicus Bres. (1912)
  • Fomes langloisii (Murrill) Sacc. & D.Sacc. (1905)
  • Fomes lazaroi Sacc. & Trotter (1925)
  • Fomes longinquus Lloyd (1925)
  • Fomes lukinsii N.Walters (1962)
  • Fomes meliae (Underw.) Murrill (1903) – British Virgin Islands
  • Fomes minimus N.Walters (1962) – Lord Howe Island
  • Fomes mirabilis C.B.Ussher (1911)
  • Fomes nigrescens Lloyd (1915)
  • Fomes nigriporus Lázaro Ibiza (1916)
  • Fomes nigrolaccatus (Cooke) Sacc. (1888)
  • Fomes niveus (Lázaro Ibiza) Sacc. & Trotter (1925)
  • Fomes noscius Corner (1932)
  • Fomes obesus (Pat.) Sacc. & Trotter (1912)
  • Fomes ostricolor Lloyd (1915)
  • Fomes pachyderma Bres. (1912)
  • Fomes perelegans Rick (1960)
  • Fomes pini-canadensis (Schwein.) Cooke (1885)
  • Fomes praerimosus (Murrill) Sacc. & D.Sacc. (1905)
  • Fomes pseudoconchatus Henn. (1908)
  • Fomes pseudosenex (Murrill) Sacc. & Trotter (1912) – Mauritius
  • Fomes pusilla Lloyd (1915)
  • Fomes rechingeri Bres. (1910)
  • Fomes silveirae Torrend (1909)
  • Fomes subamboinensis Henn. (1904)
  • Fomes subendothejus Bres. (1910)
  • Fomes subzonatus (Lázaro Ibiza) Sacc. & Trotter (1925)
  • Fomes unciatus Bres. (1922)
  • Fomes uncinatus Bres. (1922)

References
  1. Orr, D.B.; Orr, R.T. (1980). Mushrooms of Western North America. California Natural History Guides. Berkeley, California: University of California Press. p. 78. ISBN 978-0-520-03660-4.
  2. Donk, M.A. (1960). "The generic names proposed for Polyporaceae". Persoonia. 1 (2): 173–302.
  3. Fries, E.M. (1849). Summa vegetabilium Scandinaviae (in Latin). 2. p. 319.
  4. Ryvarden, Leif (2014). Poroid Fungi of Europe. Synopsis Fungorum. 31. Oslo, Norway: Fungiflora. p. 177. ISBN 978-8290724462.
  5. Kirk, P.M. "Species Fungorum (version 29th May 2017). In: Species 2000 & ITIS Catalogue of Life". Retrieved 2017-06-12.
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