Horelophus walkeri

Horelophus walkeri

This species was first described in 1913 by Armand d'Orchymont[2] from two specimens collected in Reefton, New Zealand.[3] The beetle is 2.1–3.1 mm long with a glossy brown to black body. The pronotum and elytra have translucent pale brown margins. Its small eyes are pale and protrude from its head. Its head, pronotum and elytra are covered in small and shallow punctures. The elytra have a distinctive depression. This single species, due to its primitive characteristics, is placed in its own subfamily of Horelophinae.[4]

Specimens have been collected from waterfalls at sunny sites in Nelson, Buller and Marlborough. Sites in Marlborough include Dead Horse Creek, near Canvastown and tributaries of the Pelorus River, near Pelorus Bridge. In Nelson, this beetle has been collected from Deepwater Creek, near Lyell and catchments flowing into the Wangapeka and Owen Rivers. All locations were waterways that were fast flowing, clear and cool. The riverbeds consisted of fine sedimentary rocks. The locations mostly have open canopies among beech forest. Horelophus walkeri can be found on wet rocks or hiding within moss, crevices and shaded areas within the spray zone of a waterfall. Commonly, it is found walking on wet rocks, rather than submerged like water scavenger beetles that live in similar habitats.[4]

This species hides within moss and crevices in the splash zones of small waterfalls. It is likely that they feed on algae on the smooth and wet rock surfaces in these locations. Whilst adult water scavenger beetles are normally herbivores or scavengers, larvae are carnivorous.[3] Considering that they are not found at sites with closed canopies, it is likely that they use polarised light, reflecting from the surfaces of rocks under waterfalls for finding suitable locations. Their long wings indicate that they are capable of flight.[4]

Few collections had been made of this species until 2012.[1] Surveys for this species may assist in understanding its specific habitat requirements. Currently, it has not been located in any river catchments in Nelson City, indicating that populations may not be sustained in urban environments.[4] In 2012, Horelophus walkeri was added to the list of nationally endangered beetles in New Zealand.