Hymenoptera paleobiota of Burmese amber

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018;[1] its supplement Ross 2019b covers most of 2019.[2]

Hymenoptera

Taxon Authority Year described Notes image

Abropelecinus tytthus[3]

Guo et al.

2016

A pelecinid wasp.

Alivespa colossa[4]

Wu et al.

2020

A priorvespine vespid wasp.

Alivespa hirta[4]

Wu et al.

2020

A priorvespine vespid wasp.

Ampulicomorpha janzeni[5]

Olmi et al.

2014

An embolemid wasp.

Apodolichurus diaphanus[6]

Antropov

2000

A digger wasp.

Apodolichurus sphaerocephalus[6]

Antropov

2000

A digger wasp.

Aptenoperissus amabilis[7]

Zhang & Rasnitsyn

2017

An Aptenoperissidae wasp

Aptenoperissus burmanicus[8]

Rasnitsyn & Poinar

2016

An Aptenoperissidae wasp

Aptenoperissus delicatus[7]

Zhang & Rasnitsyn

2017

An Aptenoperissidae wasp

Aptenoperissus etius[9]

Rasnitsyn & Öhm-Kühnle

2018

An Aptenoperissidae wasp

Aptenoperissus formosus[7]

Zhang & Rasnitsyn

2017

An Aptenoperissidae wasp

Aptenoperissus magnifemoris[9]

Rasnitsyn & Öhm-Kühnle

2018

An Aptenoperissidae wasp

Aptenoperissus pusillus[9]

Rasnitsyn & Öhm-Kühnle

2018

An Aptenoperissidae wasp

Aptenoperissus zonalis[10]

Zhang & Rasnitsyn

2018

An Aptenoperissidae wasp

Aquilomyrmex huangi[11]

Perrichot, Wang & Barden

2020

A haidomyrmecine ant

Archaeoserphites engeli[12]

Rasnitsyn & Öhm-Kühnle

2020

An archaeoserphitid wasp.

Archaeovespa cretacea[13]

Wu et al.

2020

A vespid wasp.

Archaeovespa engeli[13]

Wu et al.

2020

A vespid wasp.

Archaeovespa malleata[13]

Wu et al.

2020

A vespid wasp.

Archaeromma gibsoni[14]

Engel & Grimaldi

2007

A false fairy wasp.

Archeofoenus engeli[15]

Turrisi & Ellenberger

2019

A hyptiogastritine aulacid wasp.

Archeofoenus tartaricus[16]

Engel

2017

A hyptiogastritine aulacid wasp.

Archaeorhyssalus subsolanus[17]

Engel in Engel & Wang

2016

A Protorhyssalinae braconid wasp.

Archaeoteleia astropulvis[18]

Talamas in Talamas et al.

2016

A platygastroid wasp of uncertain placement.

Aureobythus decoloratus[19]

Melo & Lucena

2019

A chrysobythid chrysidoid wasp

Aureobythus punctatus[19]

Melo & Lucena

2019

A chrysobythid chrysidoid wasp

Aureobythus villosus[19]

Melo & Lucena

2019

A chrysobythid chrysidoid wasp

Auricleptes nebulosus[20]

Lucena & Melo

2018

A cuckoo wasp..

Azanichrum pilosum[20]

Lucena & Melo

2018

A cuckoo wasp.

Baeomorpha liorum[21]

Huber, Shih & Ren

2019

A rotoitid wasp.

Bethylitella cylindrella[22]

Cockerell

1917

A Bethylidae wasp

Bethylochrysis clypeata[19]

Melo & Lucena

2019

A chrysobythid chrysidoid wasp.

Bohartiura glabrata[20]

Lucena & Melo

2018

A cuckoo wasp.

Brachypelecinus euthyntus[3]

Guo et al.

2016

A pelecinid wasp.

Bryopompilus interfector[23]

Engle & Grimaldi

2006

A bryopompilid wasp.[24]

Burmadryinus cenomanianus[25]

Olmi, Xu & Guglielmino

2014

A dryinid wasp.

Burmaevania aequalis[26]

Shih et al.

2020

An evaniid wasp.

Burmaevania brevis[26]

Shih et al.

2020

An evaniid wasp.

Burmaimetsha concava[27]

Perrichot

2013

A maimetshid wasp.

Burmanteon olmii[28]

Engel

2003

An anteonine dryinid wasp

Burmaphron prolatum[29]

Engel & Grimaldi

2009

A stigmaphronid wasp

Burmaphron tridentatum[29]

Engel & Grimaldi

2009

A stigmaphronid wasp

Burmasclerogibba aptera[30]

Perkovsky et al.

2020

A member of the family Sclerogibbidae.

Burmasega ammirabilis[20]

Lucena & Melo

2018

A cuckoo wasp.

Burmasphex pilosus[31]

Melo & Rosa

2018

An Angarosphecidae wasp.

Burmasphex sulcatus[31]

Melo & Rosa

2018

An Angarosphecidae wasp.

Burmastatus triangularis[6]

Antropov

2000

A Crabronid wasp

Burminata caputaeria[32]

Haas, Burks & Krogmann

2018

A diversinitid chalcid wasp.

Burmomyrma rossi[33]

Dlussky

1996

A Falsiformicid wasp,
originally placed in Formicidae

Burmorussus mirabilis[34]

Zhang et al.

2020

A parasitic wasp belonging to the family Burmorussidae.

Burmusculus fuxii[35]

Zhang & Rasnitsyn in Zhang, Rasnitsyn & Zhang

2018

A burmusculid wasp.

Burmusculus magnus[36]

Li et al.

2020

A burmusculid wasp.

Burmusculus nuwae[35]

Zhang & Rasnitsyn in Zhang, Rasnitsyn & Zhang

2018

A burmusculid wasp.

Burmusculus shennongii[35]

Zhang & Rasnitsyn in Zhang, Rasnitsyn & Zhang

2018

A burmusculid wasp.

Caloichneumon perrarus[37]

Li et al.

2017

An ichneumon wasp.

Camelomecia janovitzi[38]

Barden & Grimaldi

2016

A stem group ant of uncertain affinity

Camelosphecia fossor[39]

Boudinot, Perrichot & Chaul

2020

A member of Formicoidea closely related to ants.

Camelosphecia venator[39]

Boudinot, Perrichot & Chaul

2020

A member of Formicoidea closely related to ants.

Cascoscelio incassa[40]

Poinar

2012

A platygastrid wasp

Ceratomyrmex ellenbergeri[11]

Perrichot, Bo, & Engel,

2016

A haidomyrmecine ant

Ceratomyrmex planus[41]

Lattke & Melo

2020

A haidomyrmecine ant

Chonidris insolita[11]

Perrichot, Wang & Barden

2020

A haidomyrmecine ant

Chrysobythus areolatus[19]

Melo & Lucena

2019

A chrysobythid chrysidoid wasp.

Cirrosphex admirabilis[6]

Antropov

2000

A wasp of uncertain placement.

Colmepsiterona cumcarena[42]

Cockx & McKellar

2018

A Pemphredoninae crabronid wasp.

Cretampulex gracilis[6]

Antropov

2000

A digger wasp.

Cretapristocera longiscapa[43]

Jouault et al.

2020

A bethylid wasp.

Cretevania bechlyi[44]

Jennings, Krogmann & Mew

2013

An evaniid wasp.

Cretevania tenuis[45]

Li et al.

2018

An ensign wasp.

Cretolixon alatum[46]

Lohrmann in Lohrmann et al.

2020

A rhopalosomatid wasp.

Cretosapyga resinicola[47]

Bennett & Engel

2005

A sapygid wasp

Cretosclerogibba antennalis[30]

Perkovsky et al.

2020

A member of the family Sclerogibbidae.

Cretosclerogibba contractocollis[30]

Perkovsky et al.

2020

A member of the family Sclerogibbidae.

Cretosclerogibba neli[30]

Perkovsky et al.

2020

A member of the family Sclerogibbidae.

Cretosclerogibba rasnitsyni[30]

Perkovsky et al.

2020

A member of the family Sclerogibbidae.

Cretospilomena familiaris[6]

Antropov

2000

A Crabronid wasp

Curiosivespa striata[48]

Perrard & Carpenter

2017

A vespid wasp.

Curiosivespa zigrasi[48]

Perrard & Carpenter

2017

A vespid wasp.

Cursoribythus silvestris[49]

Cockx & McKellar

2016

A falsiformicid wasp.

Curtevania enervia[45]

Li et al.

2018

An evanioid wasp.

Decasphex cretacicus[50]

Zheng et al.

2020

An angarosphecid wasp.

Deltoxyela engeli[51]

Wang et al.

2020

A syspastoxyelid sawfly.

Dhagnathos autokrator[11]

Perrichot, Wang & Barden

2020

A haidomyrmecine ant.

Diaspathion ortegai[52]

Engel & Huang

2015

A spathiopterygid diaprioid wasp.

Dilobops bidentata[41]

Lattke & Melo

2020

A haidomyrmecine ant

Dipteromma paradoxa[53]

Rasnitsyn et al.

2019

A mipterommatid mymarommatoid wasp.

Discoscapa apicula[54]

Poinar

2020

A discoscapid bee.

Diversinitus attenboroughi

Haas, Burks, & Krogmann

2018

A diversinitid wasp

Dryinus acutifrons[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus arakanesis[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus bamar[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus brachytarsis[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus burmensis[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus chin[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus katyae[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus laminatus[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus latitarsis[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus maderai[56]

Guglielmino et al.

2018

A dryinid wasp.

Dryinus nadezhdae[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus parisei[55]

Martynova et al.

2020

A dryinid wasp.

Dryinus taron[55]

Martynova et al.

2020

A dryinid wasp.

Edrossia vetusta[30]

Perkovsky et al.

2020

A member of the family Sclerogibbidae.

Electrofoenia jehani[57]

Jouault, Nel & Perrichot

2020

An aulacid wasp.

Electrofoenops diminuta[16]

Engel

2017

An aulacid wasp.

Electrofoenops cockerelli[15]

Turrisi & Ellenberger

2019

An aulacid wasp.

Electrofoenops rasnitsyni[15]

Turrisi & Ellenberger

2019

An Aulacid wasp.

Electrofoenus gracilipes[22]

Cockerell

1917

An Aulacidae wasp

Embolemus antiquus[58]

Perkovsky et al.

2020

An embolemid wasp.

Embolemus burmensis[58]

Perkovsky et al.

2020

An embolemid wasp.

Embolemus cretacicus[58]

Perkovsky et al.

2020

An embolemid wasp.

Embolemus ohmkuhnlei[58]

Perkovsky et al.

2020

An embolemid wasp.

Embolemus zherikhini[58]

Perkovsky et al.

2020

An embolemid wasp.

Eopelecinus inopinatus[59]

Jouault et al.

2020

A pelecinid wasp.

Eorhopalosoma gorgyra[60]

Engel

2008

A rhopalosomatid wasp.

Eorhopalosoma lohrmanni[61]

Boudinot & Dungey

2020

A rhopalosomatid wasp.

Exilaulacus loculatus[45]

Li et al.

2018

An evanioid wasp.

Exilaulacus latus[45]

Li et al.

2018

An evanioid wasp.

Foenobethylus electriphilus[22][62]

(Cockerell)

1917

A bethylid wasp.
Originally described as "Apenesia" electriphila.
Moved from "Eleganesia" electriphila.

Galloromma kachinensis[14]

Engel & Grimaldi

2007

A false fairy wasp.

Geoscelio mckellari[63]

Engel & Huang

2016

A scelionid wasp.

Gerontoformica contegus[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Gerontoformica gracilis[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Gerontoformica magnus[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Gerontoformica orientalis[64]

(Engel & Grimaldi)

2005

A sphecomyrmine ant

Gerontoformica pilosus[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Gerontoformica robustus[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Gerontoformica rugosus[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Gerontoformica spiralis[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Gerontoformica subcuspis[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Gerontoformica tendir[64]

(Barden & Grimaldi)

2014

A sphecomyrmine ant

Glabiala barbata[32]

Haas, Burks & Krogmann

2018

A diversinitid chalcid wasp.

Habraulacus zhaoi[65]

Li et al.

2015

A praeaulacid evanioid wasp.

Haidomyrmex cerberus[11]

Dlussky

1996

A haidomyrmecine ant

Haidomyrmex davidbowiei[41]

Lattke & Melo

2020

A haidomyrmecine ant

Haidomyrmex scimitarus[11]

Barden & Grimaldi

2012

A haidomyrmecine ant

Haidomyrmex zigrasi[11]

Barden & Grimaldi

2012

A haidomyrmecine ant

Heteropimpla megista[66]

Li, Shih, Kopylov & Ren in Li et al.

2019

A Labenopimplinae ichneumon wasp.

Holopsenelliscus pankowskiorum[67]

Engel

2019

A bethylid wasp.

Hybristodryinus anomalus[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus concavifrons[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus cretacicus[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus karen[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus kayin[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus konbaung[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus ligulatus[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus magnificus[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus mon[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus moutesoe[69]

Tribull, Barden & Olmi

2020

A dryinid wasp.

Hybristodryinus nalae[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus pyu[68]

Perkovsky et al.

2019

A dryinid wasp.

Hybristodryinus resinicolus[70]

Engel

2005

A dryinid wasp

Hybristodryinus shan[68]

Perkovsky et al.

2019

A dryinid wasp.

Hypselogastrion simplex[71]

Engel in Engel & Wang

2016

A gasteruptiid wasp.

Hyptiogastrites electrinus[72]

Cockerell

1917

An aulacid wasp

Kronostephanus zigrasi[73]

Engel & Grimaldi

2013

A schlettereriine crown wasp.

Lagenostephanus lii[74]

Li, Rasnitsyn, Shih & Ren

2017

A crown wasp.

Linguamyrmex brevicornis[11]

Perrichot et al

2020

A haidomyrmecine ant

Linguamyrmex rhinocerus[11]

Miao & Wang

2019

A haidomyrmecine ant

Linguamyrmex vladi[11]

Barden & Grimaldi

2017

A haidomyrmecine ant

Maimetshasia kachinensis[27]

Perrichot

2013

A maimetshid wasp.

Megalopsenella pouilloni[43]

Jouault et al.

2020

A bethylid wasp.

Melittosphex burmensis[75]

Poinar & Danforth

2006

An Apoid bee

Mendampulex monilicularis[6]

Antropov

2000

A digger wasp.

Mesevania swinhoei[76]

Basibuyuk & Rasnitsyn

2000

An ensign wasp

Miracorium tetrafoveolatum[20]

Lucena & Melo

2018

A cuckoo wasp.

Myanmarina jeannineae[77]

Li et al.

2018

A myanmarinid wasp.

Myanmarina kachin[78]

Zhang & Rasnitsyn in Zhang et al.

2018

A myanmarinid wasp.

Myanmarina lahu[78]

Zhang & Rasnitsyn in Zhang et al.

2018

A myanmarinid wasp.

Myanmarina lisu[78]

Zhang & Rasnitsyn in Zhang et al.

2018

A myanmarinid wasp.

Myanmarina sidorchukae[79]

Jouault, Rasnitsyn & Perrichot

2020

A myanmarinid wasp.

Myanmephialtites bashkuevi[80]

Jouault, Rasnitsyn & Perrichot

2020

An ohlhoffiid stephanoid wasp.

Myanmymar aresconoides[81]

Huber

2011

A fairyfly

Myanmyrma gracilis[82]

Engel & Grimaldi

2005

A sphecomyrmine ant.[39]

Myanmyrma maraudera[38][39]

Barden & Grimaldi

2016

A sphecomyrmine ant. Originally described as Gerontoformica maraudera.

Nadezhdabythus burmensis[83]

Zhang et al.

2020

A scolebythid wasp.

Newjersevania brevis[45]

Li et al.

2018

An evaniid wasp.

Newjersevania longa[45]

Li et al.

2018

An evaniid wasp.

Newjersevania rasnitsyni[26]

Shih et al.

2020

An evaniid wasp.

Novichneumon longus[37]

Li et al.

2017

An Ichneumon wasp.

Ohlhoffia robusta[80]

Jouault, Rasnitsyn & Perrichot

2020

An ohlhoffiid stephanoid wasp.

Othniodellitha mantichora[84]

Engel & Huang in Engel et al.

2016

An Evanioid wasp.

Palaeoanteon cenomanianum[85]

Perkovsky et al.

2020

A dryinid wasp.

Paleosyncrasis hongi[86]

Poinar

2019

A praeaulacid evanioid wasp.

Pangu antiquum[87]

Li et al.

2020

A panguid wasp.

Pangu yuangu[36]

Li et al.

2020

A panguid wasp.

Peleproctus dolichurus[88]

Zhang et al.

2018

A peleserphid proctotrupomorph.

Peleserphus brachyurus[88]

Zhang et al.

2018

A peleserphid proctotrupomorph.

Peleserphus genalis[88]

Zhang et al.

2018

A peleserphid proctotrupomorph.

Plumalexius ohmkuhnlei[89]

Rasnitsyn & Brothers

2020

A species of Plumalexius.

Ponomarenkoa burmensis[58]

Perkovsky et al.

2020

A dryinid wasp or an embolemid wasp.

Ponomarenkoa ellenbergeri[90]

Olmi, Xu & He

2013

A Ponomarenkoinae Dryinid wasp[90] or an embolemid wasp.[58]

Prolemistus apiformis[6]

Antropov

2000

A Crabronid wasp

Prosphex anthophilos[91]

Grimaldi, Peñalver, Barrón, et al.

2019

A probable stem-group Aculeatan of uncertain placement, possibly a panguid.[87]

Phoriostephanus exilis[92]

Engel & Huang

2016

A crown wasp.

Proterosceliopsis ambulata[93]

Talamas, Shih & Ren in Talamas et al.

2019

A proterosceliopsid platygastroid wasp.

Proterosceliopsis nigon[93]

Talamas, Shih & Ren in Talamas et al.

2019

A proterosceliopsid platygastroid wasp.

Proterosceliopsis plurima[93]

Talamas, Shih & Ren in Talamas et al.

2019

A proterosceliopsid platygastroid wasp.

Proterosceliopsis torquata[93]

Talamas, Shih & Ren in Talamas et al.

2019

A proterosceliopsid platygastroid wasp.

Proterosceliopsis wingerathi[93]

Talamas, Shih & Ren in Talamas et al.

2019

A proterosceliopsid platygastroid wasp.

Protobelyta monsirei[94]

Jouault, Perrichot & Nel

2020

A diapriid wasp.

Protobracon robusticauda[95]

Chen et al.

2020

A braconid wasp.

Protoceratomyrmex revelatus[11]

Perrichot, Wang & Barden

2020

A haidomyrmecine ant

Protofoenus swinhoei[72]

Cockerell

1917

An Aulacidae wasp

Protovespa haxairei[48]

Perrard & Carpenter

2017

A vespid wasp.

Protozigrasimecia chauli[96]

Cao, Boudinot & Gao in Cao et al.

2020

A zigrasimeciine ant.

Pseudodryinus burmensis[25]

Olmi, Xu & Guglielmino

2014

A dryinid wasp.

Pyramidibracon clypeatus[95]

Chen et al.

2020

A braconid wasp.

Raptodryinus patrickmuelleri[97]

Olmi et al

2020

A dryinid wasp

Rasnitsevania ferox[57]

Jouault, Nel & Perrichot

2020

A praeaulacid wasp.

Rhetinorhyssalus morticinus[98]

Engel

2016

A braconid wasp.

Sclerodermus quadridentatus[22]

Cockerell

1917

A Bethylidae wasp

Sclerogibba cretacica[99]

Martynova et al.

2019

A sclerogibbid wasp.

Sclerosyntexis hirsuta[100]

Wang et al.

2020

An anaxyelid sawfly.

Seneciobracon novalatus[101]

Engel & Huang in Engel et al.

2018

A braconid wasp.

Siccibythus martynovae[102]

Rasnitsyn et al.

2020

A falsiformicid wasp.

Siccibythus musculosus[49]

Cockx & McKellar

2016

A falsiformicid wasp.

Siccibythus oculatus[102]

Rasnitsyn et al.

2020

A falsiformicid wasp.

Siccibythus ohmkuhnlei[102]

Rasnitsyn et al.

2020

A falsiformicid wasp.

Siccibythus pallidus[102]

Rasnitsyn et al.

2020

A falsiformicid wasp.

Siccibythus paulus[102]

Rasnitsyn et al.

2020

A falsiformicid wasp.

Sinuevania mira[45]

Li et al.

2018

An evanioid wasp.

Sorellevania deansi[103]

Engel

2006

An ensign wasp

Striaexyela longicornis[104]

Zheng et al.

2019

A syspastoxyelid wasp.

Supraserphites draculi[105]

Rasnitsyn & Öhm-Kühnle

2019

A serphitid wasp.

Supraserphites margritae[106]

Rasnitsyn & Öhm-Kühnle

2020

A serphitid wasp.

Supraserphites sidorchukae[107]

Rasnitsyn & Öhm-Kühnle

2019

A serphitid wasp.

Supraserphites vorontsovi[106]

Rasnitsyn & Öhm-Kühnle

2020

A serphitid wasp.

Syspastoxyela pinguis[51]

Wang et al.

2020

A syspastoxyelid sawfly.

Syspastoxyela rhaphidia[108]

Engel & Huang in Engel et al.

2016

A syspastoxyelid sawfly.

Syspastoxyela simpla[51]

Wang et al.

2020

A syspastoxyelid sawfly.

Thanatotiphia nyx[109]

Engel, Ortega-Blanco & Bennett

2009

A tiphiid wasp

Tibialobracon compressicornis[95]

Chen et al.

2020

A braconid wasp.

Tichostephanus hui[110]

Engel

2019

A crown wasp.

Trigampulex pervetus[111]

(Cockerell)

1917

A wasp of uncertain placement.[6]

Uniceratops trex[62]

Colombo, Gobbi & Azevedo

2020

A bethylid wasp.

Xenodellitha preta[16]

Engel

2017

An othniodellithid evanioid wasp.

Zigrasimecia ferox[112]

Perrichot

2014

A zigrasimeciine ant

Zigrasimecia hoelldobleri[96]

Cao, Boudinot & Gao in Cao et al.

2020

A zigrasimeciine ant.

Zigrasimecia tonsora[113]

Barden & Grimaldi

2013

A Zigrasimeciine ant

Zoropelecinus periosus[3]

Guo et al.

2016

A pelecinid wasp.

Zoropelecinus zigrasi[114]

Engel & Grimaldi

2013

A pelecinid wasp.


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