List of non-avian dinosaur species preserved with evidence of feathers

Several non-avian dinosaurs were feathered. Direct evidence of feathers exists for the following species, listed in the order currently accepted evidence was first published. In all examples, the evidence described consists of feather impressions, except those genera inferred to have had feathers based on skeletal or chemical evidence, such as the presence of quill knobs (the anchor points for wing feathers on the forelimb) or a pygostyle (the fused vertebrae at the tail tip which often supports large feathers).[1]

  1. Ostromia crassipes (1970; possibly avialan[2])[3][4]
  2. Avimimus portentosus (inferred 1987: ulnar ridge)[5][6]
  3. Sinosauropteryx prima (1996)[7]
  4. Fulicopus lyellii, an ichnotaxon, possible squatting Dilophosaurus or similar (1996)[8]
  5. Protarchaeopteryx robusta (1997)[9]
  6. GMV 2124 (1997)[10]
  7. Caudipteryx zoui (1998)[11]
  8. Rahonavis ostromi (inferred 1998: quill knobs; possibly avialan[12])[13]
  9. Shuvuuia deserti (1999)[14]
  10. Beipiaosaurus inexpectus (1999)[15]
  11. Sinornithosaurus millenii (1999)[16]
  12. Caudipteryx dongi (2000)[17]
  13. Caudipteryx sp. (2000)[18]
  14. Microraptor zhaoianus (2000)[19]
  15. Nomingia gobiensis (inferred 2000: pygostyle)[20]
  16. Psittacosaurus sp.? (2002)[21]
  17. Scansoriopteryx heilmanni (2002; possibly avialan[22])[23]
  18. IVPP V13476 (2003)[24][25]
  19. Yixianosaurus longimanus (2003; possibly avialan[22])[26]
  20. Dilong paradoxus (2004)[27]
  21. Pedopenna daohugouensis (2005; possibly avialan[28])[29]
  22. Jinfengopteryx elegans (2005)[30][31]
  23. Juravenator starki (2006)[32][33]
  24. Sinocalliopteryx gigas (2007)[34]
  25. Velociraptor mongoliensis (inferred 2007: quill knobs)[35]
  26. Epidexipteryx hui (2008; possibly avialan[22])[36]
  27. Similicaudipteryx yixianensis (inferred 2008: pygostyle)[37]
  28. Anchiornis huxleyi (2009; possibly avialan[22])[38]
  29. Tianyulong confuciusi? (2009)[39]
  30. Incisivosaurus sp. (2010)[40][41]
  31. Concavenator corcovatus? (inferred 2010: quill knobs?)[42][43]
  32. Xiaotingia zhengi (2011; possibly avialan[22])[44]
  33. Yutyrannus huali (2012)[45]
  34. Sciurumimus albersdoerferi (2012)[46]
  35. Ornithomimus edmontonicus (2012)[47]
  36. Ningyuansaurus wangi (2012)[48]
  37. Eosinopteryx brevipenna (2013; possibly avialan[22])[49]
  38. Jianchangosaurus yixianensis (2013)[50]
  39. Aurornis xui (2013; possibly avialan[22])[51]
  40. Changyuraptor yangi (2014)[52]
  41. Kulindadromeus zabaikalicus? (2014)[53]
  42. Citipati osmolskae (inferred 2014: pygostyle)[54]
  43. Conchoraptor gracilis (inferred 2014: pygostyle)[54]
  44. Deinocheirus mirificus? (inferred 2014: pygostyle)[55]
  45. Yi qi (2015; possibly avialan[56])[57]
  46. Ornithomimus sp. (2015)[58]
  47. Zhenyuanlong suni (2015)[59]
  48. Dakotaraptor steini (inferred 2015: quill knobs)[60]
  49. Apatoraptor pennatus (inferred 2016: quill knobs)[61]
  50. DIP-V-15103 (2016; possibly avialan[62])[63]
  51. Jianianhualong tengi (2017)[64][65]
  52. Serikornis sungei (2017; possibly avialan[22])[66]
  53. Caihong juji (2018; possibly avialan[22])[67]
  54. Xingtianosaurus ganqi (2019)[68]
  55. Ambopteryx longibrachium (2019; possibly avialan[22])[69]
  56. Wulong bohaiensis (2020)[70]
  57. Chirostenotes pergracilis? (2020)[71]
  58. Dineobellator notohesperus (inferred 2020: quill knobs)[72]
  59. Ubirajara jubatus (2020)[73]
  • Note that the filamentous structures in some ornithischian dinosaurs (Psittacosaurus, Tianyulong and Kulindadromeus) and the pycnofibres found in some pterosaurs may or may not be homologous with the feathers of theropods.[39][74]
Fossil of Sinornithosaurus millenii, the first evidence of feathers in dromaeosaurids
Cast of a Caudipteryx fossil with feather impressions and stomach content
Fossil cast of a Sinornithosaurus millenii
Jinfengopteryx elegans fossil

References
  1. Stephen L. Brusatte, Graeme T. Lloyd, Steve C. Wang, Mark A. Norell (2014). "Gradual assembly of avian body plan culminated in rapid rates of evolution across the dinosaur-bird transition". Current Biology. 24 (20): 2386–2392. doi:10.1016/j.cub.2014.08.034. PMID 25264248.CS1 maint: uses authors parameter (link)
  2. Rauhut, Oliver WM; Tischlinger, Helmut; Foth, Christian (2019-05-14). "A non-archaeopterygid avialan theropod from the Late Jurassic of southern Germany". eLife. 8: e43789. doi:10.7554/eLife.43789. ISSN 2050-084X. PMC 6516837. PMID 31084702.
  3. Ostrom, John H. (1970). "Archaeopteryx: Notice of a "New" Specimen". Science. 170 (3957): 537–538. Bibcode:1970Sci...170..537O. doi:10.1126/science.170.3957.537. PMID 17799709. S2CID 46682559.
  4. Foth, C.; Rauhut, O.W.M. (2017). "Re-evaluation of the Haarlem Archaeopteryx and the radiation of maniraptoran theropod dinosaurs". BMC Evolutionary Biology. 17 (1): 236. doi:10.1186/s12862-017-1076-y. PMC 5712154. PMID 29197327.
  5. Kurzanov, S.M. (1987). "Avimimidae and the problem of the origin of birds." Transactions of the Joint Soviet-Mongolian Paleontological Expedition, 31: 5-92. [in Russian]
  6. Chiappe, L.M. and Witmer, L.M. (2002). Mesozoic Birds: Above the Heads of Dinosaurs. Berkeley: University of California Press, 536 pp. ISBN 0-520-20094-2
  7. Ji Q., Ji S. (1996). "On discovery of the earliest bird fossil in China and the origin of birds". Chinese Geology. 10 (233): 30–33.CS1 maint: uses authors parameter (link)
  8. Gierliński, G. (1996). "Feather-like impressions in a theropod resting trace from the Lower Jurassic of Massachusetts". Museum of Northern Arizona Bulletin. 60: 179–184.
  9. Ji, Q., and Ji, S. (1997). "A Chinese archaeopterygian, Protarchaeopteryx gen. nov." Geological Science and Technology (Di Zhi Ke Ji), 238: 38-41. Translated By Will Downs Bilby Research Center Northern Arizona University January 2001
  10. Ji, Q.; Ji, S. (1997). "Advances in Sinosauropteryx research". Chinese Geology. 7: 30–32.
  11. Currie, Philip J.; Qiang, Ji; Norell, Mark A.; Shu-An, Ji (1998). "Two feathered dinosaurs from northeastern China" (PDF). Nature. 393 (6687): 753–761. Bibcode:1998Natur.393..753Q. doi:10.1038/31635. S2CID 205001388.
  12. Agnolin, F.L.; Novas, F.E. (2011). "Unenlagiid theropods: are they members of the Dromaeosauridae (Theropoda, Maniraptora)?" (PDF). Anais da Academia Brasileira de Ciências. 83 (1): 117–162. doi:10.1590/S0001-37652011000100008. PMID 21437379. Retrieved 23 April 2011.
  13. Forster, Catherine A.; Sampson, Scott D.; Chiappe, Luis M.; Krause, David W. (1998a). "The Theropod Ancestry of Birds: New Evidence from the Late Cretaceous of Madagascar". Science. 279 (5358): 1915–1919. Bibcode:1998Sci...279.1915F. doi:10.1126/science.279.5358.1915. PMID 9506938. (HTML abstract)
  14. Schweitzer, M.H.; Watt, J.A.; Avci, R.; Knapp, L.; Chiappe, L.; Norell, M.; Marshall, M. (1999). "Beta-keratin specific immunological reactivity in feather-like structures of the Cretaceous Alvarezsaurid, Shuvuuia deserti". Journal of Experimental Zoology. 285 (2): 146–57. doi:10.1002/(SICI)1097-010X(19990815)285:2<146::AID-JEZ7>3.0.CO;2-A. PMID 10440726.
  15. Xu, Xing; Tang, Zhi-lu; Wang, Xiao-lin (1999). "A therizinosauroid dinosaur with integumentary structures from China". Nature. 399 (6734): 350–354. Bibcode:1999Natur.399..350X. doi:10.1038/20670. S2CID 204993327.
  16. Wu, Xiao-Chun; Xu, Xing; Wang, Xiao-Lin (1999). "A dromaeosaurid dinosaur with a filamentous integument from the Yixian Formation of China". Nature. 401 (6750): 262–266. Bibcode:1999Natur.401..262X. doi:10.1038/45769. S2CID 4430574.
  17. Zhou, Z.; Wang, X. (2000). "A new species of Caudipteryx from the Yixian Formation of Liaoning, northeast China" (PDF). Vertebrata PalAsiatica. 38 (2): 113–130. Archived from the original (PDF) on 2007-10-07.
  18. Zhou, Z.; Wang, X.; Zhang, F.; Xu, X. (2000). "Important features of Caudipteryx - Evidence from two nearly complete new specimens" (PDF). Vertebrata PalAsiatica. 38 (4): 241–254.
  19. Xu, Xing; Zhou, Zhonghe; Wang, Xiaolin (2000). "The smallest known non-avian theropod dinosaur" (PDF). Nature. 408 (6813): 705–708. Bibcode:2000Natur.408..705X. doi:10.1038/35047056. PMID 11130069. S2CID 4411157. Archived from the original (PDF) on 2008-12-17.
  20. Barsbold R., Osmólska H., Watabe M., Currie P.J., Tsogtbaatar K. (2000). "New Oviraptorosaur (Dinosauria, Theropoda) From Mongolia: The First Dinosaur With A Pygostyle" (PDF). Acta Palaeontologica Polonica. 45 (2): 97–106.CS1 maint: uses authors parameter (link)
  21. Mayr, Gerald; Peters, Stefan; Plodowski, Gerhard; Vogel, Olaf (2002). "Bristle-like integumentary structures at the tail of the horned dinosaur Psittacosaurus". Naturwissenschaften. 89 (8): 361–365. Bibcode:2002NW.....89..361M. doi:10.1007/s00114-002-0339-6. PMID 12435037. S2CID 17781405.
  22. Cau, Andrea (2020-02-25). "The body plan of Halszkaraptor escuilliei (Dinosauria, Theropoda) is not a transitional form along the evolution of dromaeosaurid hypercarnivory". PeerJ. 8: e8672. doi:10.7717/peerj.8672. ISSN 2167-8359. PMC 7047864. PMID 32140312.
  23. Czerkas, S.A., and Yuan, C. (2002). "An arboreal maniraptoran from northeast China." Pp. 63-95 in Czerkas, S.J. (Ed.), Feathered Dinosaurs and the Origin of Flight. The Dinosaur Museum Journal 1. The Dinosaur Museum, Blanding, U.S.A. PDF abridged version
  24. Xu, Xing; Zhou, Zhonghe; Wang, Xiaolin; Kuang, Xuewen; Zhang, Fucheng; Du, Xiangke (2003). "Four-winged dinosaurs from China". Nature. 421 (6921): 335–340. Bibcode:2003Natur.421..335X. doi:10.1038/nature01342. ISSN 0028-0836. PMID 12540892. S2CID 1160118.
  25. Xu, Xing; Li, Feng (2016). "A new microraptorine specimen (Theropoda: Dromaeosauridae) with a brief comment on the evolution of compound bones in theropods". Vertebrata PalAsiatica. 54 (4): 269–285.
  26. Xu X.; Wang X.-L. (2003). "A new maniraptoran from the Early Cretaceous Yixian Formation of western Liaoning". Vertebrata PalAsiatica. 41 (3): 195–202.
  27. Xu, X., Norell, M. A., Kuang, X., Wang, X., Zhao, Q., Jia, C. (2004). "Basal tyrannosauroids from China and evidence for protofeathers in tyrannosauroids". Nature. 431 (7009): 680–684. Bibcode:2004Natur.431..680X. doi:10.1038/nature02855. PMID 15470426. S2CID 4381777.CS1 maint: uses authors parameter (link)
  28. Hone D.W.E., Tischlinger H., Xu X., Zhang F. (2010). Farke, Andrew Allen (ed.). "The extent of the preserved feathers on the four-winged dinosaur Microraptor gui under ultraviolet light". PLOS ONE. 5 (2): e9223. Bibcode:2010PLoSO...5.9223H. doi:10.1371/journal.pone.0009223. PMC 2821398. PMID 20169153.CS1 maint: uses authors parameter (link)
  29. Xu, Xing; Zhang, Fucheng (2005). "A new maniraptoran dinosaur from China with long feathers on the metatarsus". Naturwissenschaften. 92 (4): 173–177. Bibcode:2005NW.....92..173X. doi:10.1007/s00114-004-0604-y. PMID 15685441. S2CID 789908.
  30. Ji Q., Ji S., Lu J., You H., Chen W., Liu Y., Liu Y. (2005). "First avialan bird from China (Jinfengopteryx elegans gen. et sp. nov.)". Geological Bulletin of China. 24 (3): 197–205.CS1 maint: uses authors parameter (link)
  31. Turner, Alan H.; Pol, Diego; Clarke, Julia A.; Erickson, Gregory M.; Norell, Mark A. (2007). "A basal dromaeosaurid and size evolution preceding avian flight" (PDF). Science. 317 (5843): 1378–1381. Bibcode:2007Sci...317.1378T. doi:10.1126/science.1144066. PMID 17823350. S2CID 2519726.
  32. Goehlich U.B., Tischlinger H., Chiappe L.M. (2006). "Juraventaor starki (Reptilia, Theropoda) ein nuer Raubdinosaurier aus dem Oberjura der Suedlichen Frankenalb (Sueddeutschland): Skelettanatomie und Wiechteilbefunde". Archaeopteryx. 24: 1–26.CS1 maint: uses authors parameter (link)
  33. Chiappe, Luis M.; Göhlich, Ursula B. (2010). "Anatomy of Juravenator starki (Theropoda: Coelurosauria) from the Late Jurassic of Germany" (PDF). Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 258 (3): 257–296. doi:10.1127/0077-7749/2010/0125.
  34. Ji S., Ji Q., Lu J., Yuan C. (2007). "A new giant compsognathid dinosaur with long filamentous integuments from Lower Cretaceous of Northeastern China". Acta Geologica Sinica. 81 (1): 8–15.CS1 maint: uses authors parameter (link)
  35. Turner, A.H.; Makovicky, P.J.; Norell, M.A. (2007). "Feather quill knobs in the dinosaur Velociraptor" (PDF). Science. 317 (5845): 1721. Bibcode:2007Sci...317.1721T. doi:10.1126/science.1145076. PMID 17885130. S2CID 11610649.
  36. Zhang, Fucheng; Zhou, Zhonghe; Xu, Xing; Wang, Xiaolin; Sullivan, Corwin (2008). "A bizarre Jurassic maniraptoran from China with elongate ribbon-like feathers". Nature. 455 (7216): 1105–1108. Bibcode:2008Natur.455.1105Z. doi:10.1038/nature07447. PMID 18948955. S2CID 4362560.
  37. He T., Wang X.-L., Zhou Z.-H. (2008). "A new genus and species of caudipterid dinosaur from the Lower Cretaceous Jiufotang Formation of western Liaoning, China". Vertebrata PalAsiatica. 46 (3): 178–189.CS1 maint: uses authors parameter (link)
  38. Xu, Xing; Zhao, Qi; Norell, Mark; Sullivan, Corwin; Hone, David; Erickson, Gregory; Wang, XiaoLin; Han, FengLu; Guo, Yu (2008). "A new feathered maniraptoran dinosaur fossil that fills a morphological gap in avian origin". Chinese Science Bulletin. 54 (3): 430–435. doi:10.1007/s11434-009-0009-6.
  39. Zheng, Xiao-Ting; You, Hai-Lu; Xu, Xing; Dong, Zhi-Ming (2009). "An Early Cretaceous heterodontosaurid dinosaur with filamentous integumentary structures". Nature. 458 (7236): 333–336. Bibcode:2009Natur.458..333Z. doi:10.1038/nature07856. PMID 19295609. S2CID 4423110.
  40. Xu, Xing; Zheng, Xiaoting; You, Hailu (2010). "Exceptional dinosaur fossils show ontogenetic development of early feathers". Nature. 464 (7293): 1338–1341. Bibcode:2010Natur.464.1338X. doi:10.1038/nature08965. PMID 20428169. S2CID 205220207.
  41. Xu, Xing (2020), "Filamentous Integuments in Nonavialan Theropods and Their Kin: Advances and Future Perspectives for Understanding the Evolution of Feathers", in Foth, Christian; Rauhut, Oliver W. M. (eds.), The Evolution of Feathers, Fascinating Life Sciences, Springer International Publishing, pp. 67–78, doi:10.1007/978-3-030-27223-4_5, ISBN 978-3-030-27222-7
  42. Ortega, Francisco; Escaso, Fernando; Sanz, José L. (2010). "A bizarre, humped Carcharodontosauria (Theropoda) from the Lower Cretaceous of Spain". Nature. 467 (7312): 203–206. Bibcode:2010Natur.467..203O. doi:10.1038/nature09181. PMID 20829793. S2CID 4395795.
  43. Cuesta, Elena; Ortega, Francisco; Sanz, José Luis (October 2015). "Ulnar bumps of Concavenator: Quill Knobs or Muscular scar? Myological Reconstruction of the forelimb of Concavenator corcoavtus (Lower Cretaceous, Las Hoyas, Spain)." ResearchGate.
  44. Xing Xu, Hailu You, Kai Du and Fenglu Han (28 July 2011). "An Archaeopteryx-like theropod from China and the origin of Avialae". Nature. 475 (7357): 465–470. doi:10.1038/nature10288. PMID 21796204. S2CID 205225790.CS1 maint: uses authors parameter (link)
  45. Xu, Xing; Wang, Kebai; Zhang, Ke; Ma, Qingyu; Xing, Lida; Sullivan, Corwin; Hu, Dongyu; Cheng, Shuqing; Wang, Shuo (2012). "A gigantic feathered dinosaur from the Lower Cretaceous of China". Nature. 484 (7392): 92–95. Bibcode:2012Natur.484...92X. doi:10.1038/nature10906. PMID 22481363. S2CID 29689629.
  46. Rauhut, O. W. M.; Foth, C.; Tischlinger, H.; Norell, M. A. (2012). "Exceptionally preserved juvenile megalosauroid theropod dinosaur with filamentous integument from the Late Jurassic of Germany". Proceedings of the National Academy of Sciences. 109 (29): 11746–11751. Bibcode:2012PNAS..10911746R. doi:10.1073/pnas.1203238109. PMC 3406838. PMID 22753486.
  47. Darla K. Zelenitsky, François Therrien, Gregory M. Erickson, Christopher L. DeBuhr, Yoshitsugu Kobayashi, David A. Eberth and Frank Hadfield (2012). "Feathered Non-Avian Dinosaurs from North America Provide Insight into Wing Origins". Science. 338 (6106): 510–514. Bibcode:2012Sci...338..510Z. doi:10.1126/science.1225376. PMID 23112330. S2CID 2057698.CS1 maint: uses authors parameter (link)
  48. Ji Qiang, Lü Jun-Chang, Wei Xue-Fang, Wang Xu-Ri (2012). "A new oviraptorosaur from the Yixian Formation of Jianchang, Western Liaoning Province, China". Regional Geology of China (12): 2102–2107.CS1 maint: uses authors parameter (link)
  49. Pascal Godefroit, Helena Demuynck, Gareth Dyke, Dongyu Hu, François Escuillié and Philippe Claeys (2013). "Reduced plumage and flight ability of a new Jurassic paravian theropod from China". Nature Communications. 4: Article number 1394. Bibcode:2013NatCo...4.1394G. doi:10.1038/ncomms2389. PMID 23340434.CS1 maint: uses authors parameter (link)
  50. Hanyong Pu, Yoshitsugu Kobayashi, Junchang Lü, Li Xu, Yanhua Wu, Huali Chang, Jiming Zhang, Songhai Jia (2013). "An unusual basal therizinosaur dinosaur with an ornithischian dental arrangement from northeastern China". PLOS ONE. 8 (5): Article number e63423. Bibcode:2013PLoSO...863423P. doi:10.1371/journal.pone.0063423. PMC 3667168. PMID 23734177.CS1 maint: uses authors parameter (link)
  51. Pascal Godefroit, Andrea Cau, Hu Dong-Yu, François Escuillié, Wu Wenhao, and Gareth Dyke (2013). "A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds". Nature. 498 (7454): 359–362. Bibcode:2013Natur.498..359G. doi:10.1038/nature12168. PMID 23719374. S2CID 4364892.CS1 maint: uses authors parameter (link)
  52. Gang Han, Luis M. Chiappe, Shu-An Ji, Michael Habib, Alan H. Turner, Anusuya Chinsamy, Xueling Liu, Lizhuo Han (2014). "A new raptorial dinosaur with exceptionally long feathering provides insights into dromaeosaurid flight performance". Nature Communications. 5: Article number 4382. Bibcode:2014NatCo...5.4382H. doi:10.1038/ncomms5382. PMID 25025742.CS1 maint: uses authors parameter (link)
  53. Godefroit, Pascal; Sinitsa, Sofia M.; Dhouailly, Danielle; Bolotsky, Yuri L.; Sizov, Alexander V.; McNamara, Maria E.; Benton, Michael J.; Spagna, Paul (2014). "A Jurassic ornithischian dinosaur from Siberia with both feathers and scales". Science. 25 (345): 451–455. Bibcode:2014Sci...345..451G. doi:10.1126/science.1253351. hdl:1983/a7ae6dfb-55bf-4ca4-bd8b-a5ea5f323103. PMID 25061209. S2CID 206556907.
  54. W. Scott Persons, IV, Philip J. Currie, and Mark A. Norell (2014). "Oviraptorosaur tail forms and functions". Acta Palaeontologica Polonica. doi:10.4202/app.2012.0093.CS1 maint: uses authors parameter (link)
  55. Yuong-Nam Lee, Rinchen Barsbold, Philip J. Currie, Yoshitsugu Kobayashi, Hang-Jae Lee, Pascal Godefroit, François Escuillié, and Tsogtbaatar Chinzorig (2014). "Resolving the long-standing enigmas of a giant ornithomimosaur Deinocheirus mirificus". Nature. 515 (7526): 257–260. Bibcode:2014Natur.515..257L. doi:10.1038/nature13874. PMID 25337880. S2CID 2986017.CS1 maint: uses authors parameter (link)
  56. Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (2019-07-10). "A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight". PeerJ. 7: e7247. doi:10.7717/peerj.7247. ISSN 2167-8359. PMC 6626525. PMID 31333906.
  57. Xu, X.; Zheng, X.; Sullivan, C.; Wang, X.; Xing, L.; Wang, Y.; Zhang, X.; O’Connor, J. K.; Zhang, F.; Pan, Y. (2015). "A bizarre Jurassic maniraptoran theropod with preserved evidence of membranous wings". Nature. 521 (7550): 70–3. Bibcode:2015Natur.521...70X. doi:10.1038/nature14423. PMID 25924069. S2CID 205243599.
  58. Van Der Reest, Aaron J.; Wolfe, Alexander P.; Currie, Philip J. (2016). "A densely feathered ornithomimid (Dinosauria: Theropoda) from the Upper Cretaceous Dinosaur Park Formation, Alberta, Canada". Cretaceous Research. 58: 108–117. doi:10.1016/j.cretres.2015.10.004.
  59. Lü, Junchang; Brusatte, Stephen L. (2015). "A large, short-armed, winged dromaeosaurid (Dinosauria: Theropoda) from the Early Cretaceous of China and its implications for feather evolution". Scientific Reports. 5: 11775. Bibcode:2015NatSR...511775L. doi:10.1038/srep11775. PMC 4504142. PMID 26181289.
  60. Robert A. DePalma, David A. Burnham, Larry D. Martin, Peter L. Larson, and Robert T. Bakker (2015). "The first giant raptor (Theropoda: Dromaeosauridae) from the Hell Creek Formation". Paleontological Contributions. 14: 1–15.CS1 maint: uses authors parameter (link)
  61. Gregory F. Funston; Philip J. Currie (2016). "A new caenagnathid (Dinosauria: Oviraptorosauria) from the Horseshoe Canyon Formation of Alberta, Canada, and a reevaluation of the relationships of Caenagnathidae". Journal of Vertebrate Paleontology. Online edition (4): e1160910. doi:10.1080/02724634.2016.1160910. S2CID 131090028.
  62. Rashid, Dana J.; Surya, Kevin; Chiappe, Luis M.; Carroll, Nathan; Garrett, Kimball L.; Varghese, Bino; Bailleul, Alida; O’Connor, Jingmai K.; Chapman, Susan C.; Horner, John R. (2018). "Avian tail ontogeny, pygostyle formation, and interpretation of juvenile Mesozoic specimens". Scientific Reports. 8 (1): 9014. Bibcode:2018NatSR...8.9014R. doi:10.1038/s41598-018-27336-x. ISSN 2045-2322. PMC 5997987. PMID 29899503.
  63. Xing, Lida; McKellar, Ryan C.; Xu, Xing; Li, Gang; Bai, Ming; Persons, W. Scott; Miyashita, Tetsuto; Benton, Michael J.; Zhang, Jianping; Wolfe, Alexander P.; Yi, Qiru (2016). "A Feathered Dinosaur Tail with Primitive Plumage Trapped in Mid-Cretaceous Amber". Current Biology. 26 (24): 3352–3360. doi:10.1016/j.cub.2016.10.008. PMID 27939315.
  64. Xing Xu; Philip Currie; Michael Pittman; Lida Xing; Qingjin Meng; Junchang Lü; Dongyu Hu; Congyu Yu (2017). "Mosaic evolution in an asymmetrically feathered troodontid dinosaur with transitional features". Nature Communications. 8: Article number 14972. Bibcode:2017NatCo...814972X. doi:10.1038/ncomms14972. PMC 5418581. PMID 28463233.
  65. "New Birdlike Dinosaur Had Modern Feathers". 2017-05-02. Retrieved 2017-05-03.
  66. Ulysse Lefèvre; Andrea Cau; Aude Cincotta; Dongyu Hu; Anusuya Chinsamy; François Escuillié; Pascal Godefroit (2017). "A new Jurassic theropod from China documents a transitional step in the macrostructure of feathers". The Science of Nature. 104 (9–10): Article number 74. Bibcode:2017SciNa.104...74L. doi:10.1007/s00114-017-1496-y. PMID 28831510. S2CID 32780661.
  67. Dongyu Hu; Julia A. Clarke; Chad M. Eliason; Rui Qiu; Quanguo Li; Matthew D. Shawkey; Cuilin Zhao; Liliana D’Alba; Jinkai Jiang; Xing Xu (2018). "A bony-crested Jurassic dinosaur with evidence of iridescent plumage highlights complexity in early paravian evolution". Nature Communications. 9 (1): Article number 217. Bibcode:2018NatCo...9..217H. doi:10.1038/s41467-017-02515-y. PMC 5768872. PMID 29335537.
  68. Qiu, Rui; Wang, Xiaolin; Wang, Qiang; Li, Ning; Zhang, Jialiang; Ma, Yiyun (2019). "A new caudipterid from the Lower Cretaceous of China with information on the evolution of the manus of Oviraptorosauria". Scientific Reports. 9 (1): Article number 6431. Bibcode:2019NatSR...9.6431Q. doi:10.1038/s41598-019-42547-6. PMC 6483983. PMID 31024012.
  69. Wang, Min; O'Connor, Jingmai K.; Xu, Xing; Zhou, Zhonghe (2019). "A new Jurassic scansoriopterygid and the loss of membranous wings in theropod dinosaurs". Nature. 569 (7755): 256–259. Bibcode:2019Natur.569..256W. doi:10.1038/s41586-019-1137-z. PMID 31068719. S2CID 148571099.
  70. Poust, Ashley W.; Gao, Chunling; Varricchio, David J.; Wu, Jianlin; Fengjiao, Zhang (2020). "A new microraptorine theropod from the Jehol Biota and growth in early dromaeosaurids". The Anatomical Record. 303 (4): 963–987. doi:10.1002/ar.24343. PMID 31943887. S2CID 210334980.
  71. Funston, G. F.; Currie, P. J. (2020). "New material of Chirostenotes pergracilis (Theropoda, Oviraptorosauria) from the Campanian Dinosaur Park Formation of Alberta, Canada". Historical Biology. advance online publication: 1–15. doi:10.1080/08912963.2020.1726908.
  72. Jasinski, Steven E.; Sullivan, Robert M.; Dodson, Peter (2020). "New dromaeosaurid dinosaur (Theropoda, Dromaeosauridae) from New Mexico and biodiversity of dromaeosaurids at the end of the Cretaceous". Scientific Reports. 10 (1): Article number 5105. doi:10.1038/s41598-020-61480-7. PMC 7099077. PMID 32218481.
  73. Smyth, Robert S.H.; Martill, David M.; Frey, Eberhard; Rivera-Sylva, Héctor E.; Lenz, Norbert (2020). "A maned theropod dinosaur from Gondwana with elaborate integumentary structures". Cretaceous Research: 104686. doi:10.1016/j.cretres.2020.104686.
  74. Czerkas, S.A., and Ji, Q. (2002). "A new rhamphorhynchoid with a headcrest and complex integumentary structures." Pp. 15-41 in: Czerkas, S.J. (Ed.). Feathered Dinosaurs and the Origin of Flight. Blanding, Utah: The Dinosaur Museum. ISBN 1-932075-01-1.
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