Lycopodium clavatum

Lycopodium clavatum is a spore-bearing vascular plant, growing mainly prostrate along the ground with stems up to 1 m (39 in) long; the stems are much branched, and densely clothed with small, spirally arranged microphyll leaves. The leaves are 3–5 mm long and 0.7–1 mm broad, tapered to a fine hair-like white point. The branches bearing strobili or spore cones turn erect, reaching 5–15 cm (2.0–5.9 in) above ground, and their leaves are modified as sporophylls that enclose the spore capsules or sporangia. The spore cones are yellow-green, 2–3 cm (0.79–1.18 in) long, and 5 mm (0.20 in) broad. The horizontal stems produce roots at frequent intervals along their length, allowing the stem to grow indefinitely along the ground. The stems superficially resemble small seedlings of coniferous trees, though it is not related to these.

Close-up of strobili

Lycopodium clavatum has a widespread distribution across several continents.[8][9][10][11][12][13] There are distinct subspecies and varieties in different parts of its range:

• Lycopodium clavatum subsp. clavatum
  • Lycopodium clavatum subsp. clavatum var. clavatum (Europe, Asia, North America)
  • Lycopodium clavatum subsp. clavatum var. aristatum (Mexico, Caribbean, Central America, northern South America south to northern Argentina)
  • Lycopodium clavatum subsp. clavatum var. asiaticum (Japan, northeast China)
  • Lycopodium clavatum subsp. clavatum var. borbonicum (central and southern Africa)
  • Lycopodium clavatum subsp. clavatum var. kiboanum (mountains of tropical Africa)
• Lycopodium clavatum subsp. contiguum (southern Central America, northern South America; syn. Lycopodium contiguum)

Although globally widespread, like many clubmosses, it is confined to undisturbed sites, disappearing from farmed areas and sites with regular burning. As a result, it is endangered in many areas. In the UK it is one of 101 species named as a high priority for conservation by the wild plant charity Plantlife.

Other common names

Common names for this species include wolf's-foot clubmoss, common clubmoss, wolf-paw clubmoss, running ground-pine, running pine,[7] running moss, princess pine, and others.

Use

The spores of this moss, "lycopodium powder", are explosive if present in the air in high enough densities. They were used as flash powder in early photography and magic acts.

Active constituents

Bioactive secondary metabolites in clubmosses include triterpenoids with acetylcholinesterase inhibitor activity isolated from this species.[14]

• 
  L. clavatum ssp. clavatum var. clavatum, with strobili
• 
  L. clavatum with strobili

• Lycopodium clavatum, running ground pine, Flora, fauna, earth, and sky...The natural history of the northwoods
• Bioimagesphotos
• Jepson Manual Treatment, University of California
• Calphotos Photo gallery, University of California
• photo of herbarium specimen at Missouri Botanical Garden, collected in Dominican Republic in 1967

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