Poecile

Poecile
	Willow tit, Poecile montanus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Passeriformes
Family:	Paridae
Genus:	Poecile; Kaup, 1829
Species
	see text

Poecile is a genus of birds in the tit family Paridae. It contains 15 species, which are scattered across North America, Europe and Asia; the North American species are the chickadees. In the past, most authorities retained Poecile as a subgenus within the genus Parus, but treatment as a distinct genus, initiated by the American Ornithologists Union, is now widely accepted.[1] This is supported by mtDNA cyrochrome b sequence analysis.[2]

The genus Poecile was erected by the German naturalist Johann Jakob Kaup in 1829.[3] The type species was subsequently designated as the marsh tit (Poecile palustris) by English zoologist George Robert Gray in 1842.[4][5] The name Poecile is from Ancient Greek poikilos "colourful". A related word poikilidos denoted an unidentified small bird.[6] It has traditionally been treated as feminine (giving name endings such as cincta); however, this was not specified by the original genus author Johann Jakob Kaup, and under the ICZN the genus name must therefore be treated by default as masculine, giving name endings such as cinctus.[1]

	White-browed tit (Poecile superciliosus)

	Sombre tit (Poecile lugubris)

Grey-headed chickadee (Poecile cinctus)

	Chestnut-backed chickadee (Poecile rufescens)

	Boreal chickadee (Poecile hudsonicus)

Mexican chickadee (Poecile sclateri)

Carolina chickadee (Poecile carolinensis)

	Black-capped chickadee (Poecile atricapillus)

	Mountain chickadee (Poecile gambeli)

Père David's tit (Poecile davidi)

	Black-bibbed tit (Poecile hypermelaenus)

	Marsh tit (Poecile palustris)

Sichuan tit (Poecile weigoldicus)

	Caspian tit (Poecile hyrcanus)

	Willow tit (Poecile montanus)

Phylogeny of the Poecile based on Tritsch et al. 2017.[7]

Species

The genus includes the following fifteen species:[8]

Common Name	Scientific name	Distribution
White-browed tit	Poecile superciliosus	central China and Tibet.
Sombre tit	Poecile lugubris	southeast Europe and southwest Asia
Grey-headed chickadee (North American name) or Siberian tit (European name)	Poecile cinctus	subarctic Scandinavia and northern Asia, and also into North America in Alaska and the far northwest of Canada
Chestnut-backed chickadee	Poecile rufescens	Pacific Northwest of the United States and western Canada, from southern Alaska to southwestern California
Boreal chickadee	Poecile hudsonicus	Canada, Alaska, and the northern edges of the northernmost portions of the lower forty-eight United States
Mexican chickadee	Poecile sclateri	Mexico
Carolina chickadee	Poecile carolinensis	United States from New Jersey west to southern Kansas and south to Florida and Texas
Black-capped chickadee	Poecile atricapillus	Across North America, from New England to Newfoundland in the east, and from Washington to Alaska in the west
Mountain chickadee	Poecile gambeli	western United States
Père David's tit	Poecile davidi	central China in southern Gansu, western Hubei, southern Shaanxi and Sichuan
Black-bibbed tit	Poecile hypermelaenus	central and eastern China to southeast Tibet and western Myanmar.
Marsh tit	Poecile palustris	temperate Europe and northern Asia
Sichuan tit	Poecile weigoldicus	central China
Caspian tit	Poecile hyrcanus	northern Iran, just extending into Azerbaijan.
Willow tit	Poecile montanus	temperate and subarctic Europe and northern Asia

References

Gosler, A.; Clement, P.; Bonan, A. (2019) [2007]. del Hoyo, J.; Elliott, A.; Sargatal, J.; Christie, D.A.; de Juana, E. (eds.). "Tits and Chickadees (Paridae)". Handbook of the Birds of the World Alive. Lynx Edicions. Retrieved 27 August 2019.
Gill, F.B.; Slikas, B.; Sheldon, F.H. (2005). "Phylogeny of titmice (Paridae): II. Species relationships based on sequences of the mitochondrial cytochrome-b gene". Auk. 122 (1): 121–143. doi:10.1642/0004-8038(2005)122[0121:POTPIS]2.0.CO;2.
Kaup, Johann Jakob (1829). Skizzirte Entwickelungs-Geschichte und natürliches System der europäischen Thierwelt (in German). Darmstadt: Carl Wilhelm Leske. p. 114.
Gray (1842). Appendix to a List of the Genera of Birds (2nd ed.). London: R. and J.E. Taylor. p. 8.
Dickinson, E.C.; Christidis, L., eds. (2014). The Howard & Moore Complete Checklist of the Birds of the World. Volume 2: Passerines (4th ed.). Eastbourne, UK: Aves Press. p. 428. ISBN 978-0-9568611-2-2.
Jobling, J.A. (2018). del Hoyo, J.; Elliott, A.; Sargatal, J.; Christie, D.A.; de Juana, E. (eds.). "Key to Scientific Names in Ornithology". Handbook of the Birds of the World Alive. Lynx Edicions. Retrieved 15 May 2018.
Tritsch, Christian; Martens, Jochen; Sun, Yue-Hua; Heim, Wieland; Strutzenberger, Patrick; Päckert, Martin (2017). "Improved sampling at the subspecies level solves a taxonomic dilemma – A case study of two enigmatic Chinese tit species (Aves, Passeriformes, Paridae, Poecile)". Molecular Phylogenetics and Evolution. 107: 538–550. doi:10.1016/j.ympev.2016.12.014.
Gill, Frank; Donsker, David (eds.). "Waxwings and their allies, tits & penduline tits". World Bird List Version 6.1. International Ornithologists' Union. Retrieved 15 February 2016.

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[hbw-1] Gosler, A.; Clement, P.; Bonan, A. (2019) [2007]. del Hoyo, J.; Elliott, A.; Sargatal, J.; Christie, D.A.; de Juana, E. (eds.). "Tits and Chickadees (Paridae)". Handbook of the Birds of the World Alive. Lynx Edicions. Retrieved 27 August 2019.

[2] Gill, F.B.; Slikas, B.; Sheldon, F.H. (2005). "Phylogeny of titmice (Paridae): II. Species relationships based on sequences of the mitochondrial cytochrome-b gene". Auk. 122 (1): 121–143. doi:10.1642/0004-8038(2005)122[0121:POTPIS]2.0.CO;2.

[3] Kaup, Johann Jakob (1829). Skizzirte Entwickelungs-Geschichte und natürliches System der europäischen Thierwelt (in German). Darmstadt: Carl Wilhelm Leske. p. 114.

[4] Gray (1842). Appendix to a List of the Genera of Birds (2nd ed.). London: R. and J.E. Taylor. p. 8.

[5] Dickinson, E.C.; Christidis, L., eds. (2014). The Howard & Moore Complete Checklist of the Birds of the World. Volume 2: Passerines (4th ed.). Eastbourne, UK: Aves Press. p. 428. ISBN 978-0-9568611-2-2.

[hbwkey-6] Jobling, J.A. (2018). del Hoyo, J.; Elliott, A.; Sargatal, J.; Christie, D.A.; de Juana, E. (eds.). "Key to Scientific Names in Ornithology". Handbook of the Birds of the World Alive. Lynx Edicions. Retrieved 15 May 2018.

[7] Tritsch, Christian; Martens, Jochen; Sun, Yue-Hua; Heim, Wieland; Strutzenberger, Patrick; Päckert, Martin (2017). "Improved sampling at the subspecies level solves a taxonomic dilemma – A case study of two enigmatic Chinese tit species (Aves, Passeriformes, Paridae, Poecile)". Molecular Phylogenetics and Evolution. 107: 538–550. doi:10.1016/j.ympev.2016.12.014.

[ioc-8] Gill, Frank; Donsker, David (eds.). "Waxwings and their allies, tits & penduline tits". World Bird List Version 6.1. International Ornithologists' Union. Retrieved 15 February 2016.