Urocoptidae

Urocoptidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea.[2]

Urocoptidae
Apertural view of a shell of Holospira elizabethae
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Heterobranchia
Superorder: Eupulmonata
Order: Stylommatophora
Superfamily: Urocoptoidea
Family: Urocoptidae
Pilsbry, 1898[1] (1868)

Taxonomy

2005 taxonomy

The family Urocoptidae was classified in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[3]

The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

  • Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868
  • Apomatinae Paul, 1982
  • Brachypodellinae H. B. Baker, 1956
  • Eucalodiinae P. Fischer & Crosse, 1873
  • Holospirinae Pilsbry, 1946
  • Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972
  • Tetrentodontinae Bartsch, 1943

2008 taxonomy

Uit de Weerd (2008)[2] moved the Urocoptidae to the newly established superfamily Urocoptoidea based on molecular phylogeny research.[2]

The following cladogram based on Bayesian analysis of sequences of 28S ribosomal RNA shows the phylogenic relations within the Urocoptoidea.[2] There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.[2]

Urocoptoidea
 
 
 
 
 
 

Macroceramus microdon

 
 

Microceramus pontificus

 
 
 
 

Torrecoptis holguinensis

 
 

Autocoptis menkeana

 
 
 
 
 

Simplicervix inornata

 
 

Brachypodella dominicensis gabbi

 
 
 
 
 
 
 

Dissotropis sp. nov.

 
 

Anisospira lebmanni

 
 
 

Archegocoptis crenata

 
 
 

Eucalodium sp.

 
 
 
Cerionidae

Cerion striatellum

 
 
 
 

Coelostemma sp. nov.

 
 

Hendersoniella lux lux

 
 

Holospira fortisculpta

 
 
 

Epirobia polygyra

 
 

2012 taxonomy

Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails (Epirobia, Propilsbrya, Pectinistemma, Gyrocion, Prionoloplax), that were previously placed within Urocoptidae.[4] Data from molecular phylogeny research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.[4]

Genera

Genera in the family Urocoptidae include:

Urocoptinae

  • Autocoptis Pilsbry, 1902[2]
  • Cylindrella L. Pfeiffer, 1840
  • Urocoptis Beck, 1837 - type genus of the family Urocoptidae[3]

Apominae

  • Apoma Beck, 1837 - type genus of the subfamily Apomatinae[3]
  • Simplicervix Pilsbry, 1903[2]

Brachypodellinae

  • Brachypodella Beck, 1837 - type genus of the subfamily Brachypodellinae[3]

Eucalodiinae

  • Anisospira Strebel, 1880[2]
  • Archegocoptis Pilsbry, 1903[2]
  • †Condonella McLellan, 1927[5]
  • Dissotropis Bartsch, 1906[2]
  • Eucalodium Crosse & P. Fischer, 1868 - type genus of the subfamily Eucalodiinae[3]

Holospirinae

  • Coelostemma Dall, 1895[2]
  • Hendersoniella Dall, 1905[2]
  • Holospira Martens, 1860 - type genus of the subfamily Holospirinae[3]

Microceraminae

  • Microceramus Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae[3]
  • Macroceramus Guilding, 1828[2]

Tetrentodontinae

  • Tetrentodon Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae[3]
  • Torrecoptis Bartsch, 1943[2]

Other genera include:[6]

  • Allocoptis Thompson & Franz, 1976
  • Amphicosmia Pilsbry & Vanatta, 1898
  • Anoma
  • Arangia
  • Bactrocoptis
  • Badiofax
  • Bostrichocentrum
  • Bostrychocentrum
  • Callonia
  • Centralia
  • Cochlodinella
  • Coelocentrum
  • Geoscala
  • Gongylostoma
  • Gyraxis
  • Haplocion
  • Heterocoptis
  • Idiostemma
  • Liocallonia
  • Metastoma
  • Mychostoma
  • Nesocoptis
  • Nodulia
  • Oligostylus
  • Organocoptis
  • Pfeiffericoptis
  • Pineria
  • Pleurostemma
  • Poeycoptis
  • Pseudopineria
  • Pycnoptychia Pilsbry & Vanatta, 1898
  • Sagracoptis
  • Spiroceramus
  • Spirostemma
  • Strophina
  • Tomelasmus
  • Trilamellaxis
  • Uncinicoptis

References
  1. Pilsbry H. A. (1898). The Nautilus 11(9): 107.
  2. Uit De Weerd, D. R. (2008). "Delimitation and phylogenetics of the diverse land-snail family Urocoptidae (Gastropoda: Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies. 74 (4): 317–29. doi:10.1093/mollus/eyn023.
  3. Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
  4. Thompson F. G. (2012). "The land snail genus Epirobia and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". Bulletin Florida Museum of Natural History 51: 167-215. PDF.
  5. Roth, B. (2000). "Upper Cretaceous (Campanian) land snails (Gastropoda: Stylommatophora) from Washington and California". Journal of Molluscan Studies. 66 (3): 373–381. doi:10.1093/mollus/66.3.373.
  6. http://pick4.pick.uga.edu/mp/20q?search=Urocoptidae

Further reading
  • Bartsch, P. (1943). "Notes on Mexican Urocoptid mollusks". Journal of the Washington Academy of Sciences. 33 (2): 54–59.; 7 figs.
  • Carlos de la Torre. 1912. New Cuban Urocoptis of the U. cinerea group. The Nautilus, volume 26, number 4, pages 55-58, plate 6.
  • Kabat A. R., Herschler R. & González G. A. (2012). "Resolution of taxonomic problems associated with the complex publication history of the seminal Torre and Bartsch monograph on Cuban Urocoptidae (Gastropoda, Pulmonata)". Zootaxa 3362: 43-53. PDF.
  • Pilsbry H. A. & Henderson J. B. jr. (February 1913). Two new Cuban Urocoptidae. The Nautilus, volume 26, number 10, pages 109-110.
  • (in Spanish) Miguel L. Jaume & Alfredo de la Torre. 1976. Los Urocoptidae de Cuba (Mollusca-Pulmonata). Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp.
  • Mesa Garcia, R; Jaume, ML (1982). "Zoogeography, ecology, and protection of the Urocoptidae family in Cuba (Gastropoda-Pulmonata). II". Revista Cubana de Medicina Tropical. 34 (2): 172–92. PMID 6763741.
  • Richardson C. L. 1991. Urocoptidae: catalogue of species. Tryonia 22: 1-245.
  • Tryon G. W. (1903) Manual of Conchology. Second series: Pulmonata. Volume 15. Urocoptidae. Continued by H. A. Pilsbry.
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