Frullania

Frullania is the only genus of liverworts in family Frullaniaceae. It contains the following species:

Frullania
Temporal range: Cenomanian–Present
Frullania tamarisci
Scientific classification
Kingdom:
Division:
Class:
Order:
Family:
Frullaniaceae
Genus:
Frullania

Raddi
Species

See text

Frullania fragilifolia

A

  • Frullania aculeata Taylor, 1846
  • Frullania acutata Caspary, 1887
  • Frullania acutiloba Gerola, 1947
  • Frullania akiyamae Hattori, 1986
  • Frullania albertii Stephani, 1916
  • Frullania allanii Hodgson, 1949
  • Frullania allionii Stephani, 1910
  • Frullania alpina Stephani, 1911
  • Frullania alstonii Verdoorn, 1930
  • Frullania alstonii var. pfleidereri Hattori, 1972
  • Frullania alternans Nees In G., L. & N., 1845
  • Frullania amamiensis Kamimura, 1968
  • Frullania ambronnii Stephani, 1916
  • Frullania amplicrania Stephani, 1910
  • Frullania ampullifera Jack & Stephani In Stephani, 1894
  • Frullania anderssonii Ångström, 1873
  • Frullania angulata Mitten, 1863
  • Frullania angulata F. Serratoides Vanden Berghen, 1983
  • Frullania angulata var. laciniata Demaret & Vanden Berghen, 1950
  • Frullania angustistipa Stephani, 1908
  • Frullania anomala Hodgson, 1949
  • Frullania antaresensis Hattori, 1980
  • Frullania aoshimensis Horikawa, 1929
  • Frullania apertilobula Gerola, 1947
  • Frullania apicalis Mitten, 1879
  • Frullania apicalis var. camerunensis Vanden Berghen, 1976
  • Frullania apiculata (R., B. & N.) Nees In G., L. & N., 1845
  • Frullania apiculata var. goebelii Schiffner, 1893
  • Frullania apiculata var. guianensis Lindenberg & Gottsche, 1851
  • Frullania apiculata var. laxa Nees In G., L. & N., 1845
  • Frullania apollinarii Stephani, 1911
  • Frullania aposinensis Hattori & Lin, 1985
  • Frullania appalachiana Schuster, 1983
  • Frullania appendistipula Hattori, 1972
  • Frullania appendistipula var. spinifera Hattori, 1974
  • Frullania arecae (Sprengel) Gottsche, 1863
  • Frullania arecae var. spiniloba (Stephani) Yuzawa, 1991
  • Frullania armata Herzog & Clark In Clark, 1954
  • Frullania armatifolia Verdoorn, 1932
  • Frullania armitiana Stephani, 1911
  • Frullania armitiana var. inflexula Hattori, 1988
  • Frullania armitiana var. longe-Attenuata (Hattori) Hattori, 1980
  • Frullania arsenii Stephani, 1924
  • Frullania asagrayana Montagne, 1842
  • Frullania astrolabea Stephani, 1910
  • Frullania aterrima (Hooker & Taylor) Hooker & Taylor In G., L. & N., 1845
  • Frullania aterrima var. lepida Hodgson, 1949
  • Frullania atrata (Swartz) Montagne, 1839
  • Frullania atrata var. flaccida (Weber) Nees In G., L. & N., 1845
  • Frullania atrata var. mexicana Nees In G., L. & N., 1845
  • Frullania atrosanguinea Taylor Ex Spruce, 1884
  • Frullania attenuata Stephani, 1911
  • Frullania auriculata Hattori, 1985
  • Frullania azorica Sim-Sim, Sergio, Mues & Kraut, 1995

B

  • Frullania baileyana Stephani, 1910
  • Frullania bakeri Stephani, 1916
  • Frullania baladina Gottsche Ex Stephani, 1894
  • Frullania baladina var. edentata Hattori, 1984
  • Frullania baltica Grolle, 1985
  • Frullania baumannii Hattori, 1977
  • Frullania beauverdii Stephani, 1916
  • Frullania bella Stephani, 1911
  • Frullania belmorensis Stephani In Stephani & Watts, 1914
  • Frullania benjaminiana Inoue In Hattori, 1975
  • Frullania bergmanii Hattori, 1974
  • Frullania berthoumieui Stephani, 1894
  • Frullania beyrichiana (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
  • Frullania bhutanensis Hattori, 1971
  • Frullania bicornistipula Spruce, 1884
  • Frullania blastopetala Hattori, 1984
  • Frullania blepharozia Spruce, 1884
  • Frullania bogotensis Stephani, 1910
  • Frullania bolanderi Austin, 1869
  • Frullania bonariensis Reiner, 1988
  • Frullania bonincola Hattori, 1978
  • Frullania borbonica Lindenberg In G., L. & N., 1845
  • Frullania boveana Massalongo, 1885
  • Frullania brachycarpa Spruce, 1889
  • Frullania brasiliensis Raddi, 1822
  • Frullania brasiliensis var. cylindrica (Gottsche In Lehmann) Spruce, 1884
  • Frullania brasiliensis var. cylindrica F. Lindigii (Gottsche) Spruce, 1884
  • Frullania breuteliana Gottsche In G., L. & N., 1845
  • Frullania brevicalycina Stephani, 1894
  • Frullania brittoniae Evans, 1897
  • Frullania brotheri Stephani, 1894
  • Frullania brunnea (Sprengel) Drège, 1843
  • Frullania buchtienii Herzog, 1942
  • Frullania bullata Stephani, 1910

C

  • Frullania caduca Hattori, 1980
  • Frullania caespitans Beauverd In Stephani, 1924
  • Frullania caffraria Stephani, 1894
  • Frullania caffraria F. Anomala Vanden Berghen, 1976
  • Frullania calcarata Ångström, 1873
  • Frullania caldensis Ångström, 1876
  • Frullania caledonica Gottsche Ex Stephani, 1894
  • Frullania californica (Austin Ex Underwood) Evans, 1897
  • Frullania campanulata Sande Lacoste, 1853
  • Frullania campanulata var. caduca Verdoorn, 1930
  • Frullania campanulata var. malesiaca (Verdoorn) Hattori, 1975
  • Frullania canaliculata Gottsche Ex Stephani, 1910
  • Frullania capensis Gottsche In G., L. & N., 1845
  • Frullania capillaris Stephani, 1911
  • Frullania carrii Kamimura & Hattori In Hattori & Kamimura, 1973
  • Frullania casparyi Grolle, 1985
  • Frullania cataractarum Stephani, 1911
  • Frullania caulisequa (Nees In Martius) Montagne, 1839
  • Frullania cavallii Gola, 1907
  • Frullania cesatiana De Notaris, 1865
  • Frullania changii Hattori & Gao, 1985
  • Frullania chenii Hattori & Lin, 1985
  • Frullania chevalieri (Schuster) Schuster, 1992
  • Frullania chiapasana Stephani, 1910
  • Frullania chilcootiensis Stephani, 1887
  • Frullania chilensis Stephani, 1894
  • Frullania chiovendae Gola, 1914
  • Frullania chodatii Beauverd In Stephani, 1924
  • Frullania ciliata Lindenberg & Gottsche In G., L. & N., 1847
  • Frullania cinchonae Gottsche In G., L. & N., 1845
  • Frullania clandestina (Nees & Montagne) Nees In G., L. & N., 1845
  • Frullania clavata (Hooker & Taylor) Taylor In G., L. & N., 1845
  • Frullania claviloba Stephani, 1911
  • Frullania clemensiana Verdoorn, 1932
  • Frullania cobrensis Gottsche Ex Stephani, 1894
  • Frullania compacta Gottsche Ex Stephani, 1911
  • Frullania complicata Stephani, 1911
  • Frullania confertiloba Stephani, 1910
  • Frullania consociata Stephani, 1910
  • Frullania contracta Stephani, 1911
  • Frullania controversa Beauverd In Stephani, 1924
  • Frullania convoluta Lindenberg & Hampe In Hampe, 1851
  • Frullania convoluta var. ampliata Herzog, 1953
  • Frullania cordaeana Lindenberg In G., L. & N., 1845
  • Frullania cordistipula (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
  • Frullania cordistipula var. (Beta) mutica Nees In G., L. & N., 1845
  • Frullania cordistipula var. (Gamma) regularis Nees In G., L. & N., 1845
  • Frullania cordistipula var. dentistipula Hattori, 1986
  • Frullania cornuta Stephani, 1911
  • Frullania crassitexta Stephani, 1910
  • Frullania crawfordii Stephani, 1894
  • Frullania crenulifolia Jack & Stephani, 1892
  • Frullania crinoidea Spruce Ex Stephani In Stephani, 1911
  • Frullania crispiloba Stephani, 1894
  • Frullania crispiplicata Yuzawa & Hattori Ex Konstantinova, Potemkin & Schijakov, 1992
  • Frullania cristata Hattori, 1981
  • Frullania cuencensis Taylor, 1846
  • Frullania cuneatistipula Stephani, 1924
  • Frullania cuneiloba Nees In G., L. & N., 1845
  • Frullania curviramea Stephani, 1911
  • Frullania curvirostris Colenso, 1889
  • Frullania curvistipula Stephani, 1911
  • Frullania curvistipula var. falcatidenta Hattori, 1982
  • Frullania curvistipula var. lamii Verdoorn, 1930
  • Frullania curvistipula var. latistipula Hattori, 1978
  • Frullania cuspidifolia Stephani, 1911
  • Frullania cuspiloba Stephani, 1910
  • Frullania cyparioides (Schwägrichen) Schwaegrichen In G., L. & N., 1845

D

  • Frullania darwinii Gradstein & Uribe In Uribe, 2004
  • Frullania davurica Hampe In G., L. & N., 1845
  • Frullania davurica F. Dorsoblastos (Hattori) Hattori & Lin, 1985
  • Frullania davurica F. Microphylla (Massalongo) Hattori & Lin, 1985
  • Frullania davurica var. chichibuensis (Kamimura) Hattori, 1976
  • Frullania davurica var. concava Chang In Gao & Chang, 1981
  • Frullania debilis Stephani Ex Hattori, 1974
  • Frullania decidua Spruce, 1884
  • Frullania deflexa Mitten, 1862
  • Frullania degelii Arnell, 1959
  • Frullania densifolia Subsp. Andamana Hattori Ex Singh In Nath & Asthana, 2001
  • Frullania densiloba Stephani Ex Evans, 1906
  • Frullania dentata Hattori, 1974
  • Frullania dentata var. secernens Hattori, 1988
  • Frullania dentifera Hattori & Streimann, 1985
  • Frullania dentiloba Hattori, 1975
  • Frullania deplanata Mitten In Hooker, 1855
  • Frullania deppii Gottsche Ex Lehmann, 1844
  • Frullania depressa Mitten, 1863
  • Frullania dilatata Subsp. (Dilatata 1982) (L.) Dumortier, 1835
  • Frullania dilatata Subsp. (Dilatata 1982) F. Fuscovirens Jorgensen, 1934
  • Frullania dilatata Subsp. (Dilatata 1982) var. anomala Corbiere, 1889
  • Frullania dilatata Subsp. (Dilatata 1982) var. macrotus Nees, 1838
  • Frullania dilatata Subsp. (Dilatata 1982) var. subtilissima Nees, 1838
  • Frullania dilatata Subsp. Asiatica Hattori, 1982
  • Frullania diptera (Lehmann & Lindenberg) Drège, 1843
  • Frullania dispar Nees In G., L. & N., 1845
  • Frullania diversitexta Stephani, 1897
  • Frullania donnellii Austin, 1879
  • Frullania dulimensis Uribe, 2006
  • Frullania durifolia Stephani, 1894
  • Frullania dusenii Stephani In Dusen, 1905
  • Frullania duthiana Stephani, 1910
  • Frullania duthiana var. laevis Hattori In Ohashi, 1975
  • Frullania duthiana var. szechuanensis Hattori & Gao, 1985

E

  • Frullania eboracensis (Subsp. Eboracensis 1992) Gottsche Ex Lehmann, 1844
  • Frullania eboracensis Subsp. Virginica (Gottsche In Lehmann) Schuster, 1992
  • Frullania echinantha Hattori, 1974
  • Frullania echinatella Hattori, 1988
  • Frullania ecklonii (Sprengel) Sprengel In G., L. & N., 1845
  • Frullania ecklonii F. Robustior Gottsche, 1863
  • Frullania ecklonii F. Tenerior Sprengel, 1847
  • Frullania ecklonii var. huitamalcensis Gottsche, 1863
  • Frullania ecklonii var. rufescens Sprengel, 1847
  • Frullania ecuadoriensis Stephani, 1911
  • Frullania elegans Lehmann, 1857
  • Frullania elephantum Hattori, 1977
  • Frullania engelii Hattori, 1983
  • Frullania epiphylla Subsp. Fijiensis Hattori, 1985
  • Frullania eplicata Stephani, 1911
  • Frullania ericoides (Nees In Martius) Montagne, 1839
  • Frullania ericoides F. Ericoides (Nees) Verdoorn Ex Piippo, 1990
  • Frullania ericoides var. planescens (Verdoorn) Hattori, 1984
  • Frullania erostrata Hattori, 1974
  • Frullania errans Verdoorn, 1930
  • Frullania errans var. angulistipula Hattori, 1972
  • Frullania esenbeckiana Beauverd Ex Stephani, 1924
  • Frullania evelynae Hattori & Thaithong, 1978
  • Frullania evoluta Mitten, 1861
  • Frullania expansa Stephani, 1897
  • Frullania eymae Hattori, 1975
  • Frullania eymae var. crispidentata Hattori & Streimann, 1985

F

  • Frullania falciloba Taylor Ex Lehmann, 1844
  • Frullania fallax Gottsche In G., L. & N., 1845
  • Frullania falsicornuta Hattori, 1986
  • Frullania falsisinuata Hattori & Piippo, 1986
  • Frullania falsisinuata F. Parvistylata (Hattori) Hattori & Piippo, 1986
  • Frullania falsisinuata var. crispidentata Hattori & Piippo, 1986
  • Frullania fauriana Stephani, 1894
  • Frullania fauriana F. Emarginata Kamimura, 1952
  • Frullania fengyangshanensis Zhu & So, 1997
  • Frullania ferdinandi-Muelleri Stephani, 1910
  • Frullania fertilis var. major Massalongo, 1885
  • Frullania flammea Taylor Ex Spruce, 1884
  • Frullania flexicaulis Spruce, 1884
  • Frullania flexuosa Hattori, 1983
  • Frullania fragilifolia (Taylor) Taylor In G., L. & N., 1845
  • Frullania franciscana Howe, 1894
  • Frullania fuegiana Stephani, 1910
  • Frullania fugax (Hooker & Taylor) Taylor In G., L. & N., 1845
  • Frullania fulfordiae Hattori, 1987
  • Frullania fusco-Virens Stephani, 1910
  • Frullania fusco-Virens var. gemmipara (Schuster & Hattori In Hattori) Hattori & Lin, 1985

G

  • Frullania gabonensis Vanden Berghen, 1976
  • Frullania gaoligongensis Bai & Gao, 1999
  • Frullania gaudichaudii (Nees & Montagne) Nees & Montagne In G., L. & N., 1845
  • Frullania gaudichaudii F. Hasseltii (Sande Lacoste) Verdoorn, 1930
  • Frullania gemmulosa Hattori & Thaithong In Hattori, Thaithong & Kitagawa, 1977
  • Frullania gibbosa (Nees) Nees In Montagne, 1840
  • Frullania gibbosa var. major Gottsche, 1863
  • Frullania gigantea Stephani, 1911
  • Frullania giraldiana Massalongo, 1897
  • Frullania giraldiana var. handelii (Verdoorn) Hattori, 1972
  • Frullania globosa Hattori & Streimann, 1985
  • Frullania glomerata (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg Ex Montagne, 1838
  • Frullania gracilicaulis Hattori, 1977
  • Frullania gracilis Subsp. (Gracilis 1978) (Reinwardt, Blume & Nees) Gottsche In G., L. & N., 1845
  • Frullania gracilis Subsp. (Gracilis 1978) var. brevior Nees In G., L. & N., 1845
  • Frullania gracilis Subsp. (Gracilis 1978) var. vittata Hattori, 1986
  • Frullania gracilis Subsp. Zennoskei Hattori & Thaithong, 1978
  • Frullania gradsteinii Yuzawa, Mues & Hattori, 1987
  • Frullania granatensis Gottsche, 1864
  • Frullania grandifolia Stephani, 1911
  • Frullania grandilobula Hattori & Piippo, 1986
  • Frullania grandistipula Lindenberg In G., L. & N., 1845
  • Frullania griffithsiana Gottsche In G., L. & N., 1846
  • Frullania grolleana Hattori, 1972
  • Frullania grossiclava Stephani, 1910
  • Frullania grossifolia Stephani, 1911
  • Frullania guadalupensis Gottsche Ex Stephani, 1911
  • Frullania gualaquizana Stephani, 1911
  • Frullania guatemalensis Stephani, 1911

H

  • Frullania haeckeriana Lindenberg In G., L. & N., 1846
  • Frullania haematocysta Spruce, 1884
  • Frullania hainanensis Hattori & Lin, 1986
  • Frullania hamata Stephani, 1911
  • Frullania hamaticoma Stephani, 1889
  • Frullania hamatiloba Stephani, 1910
  • Frullania hamatiloba F. Grosse-Appendiculata Hattori, 1944
  • Frullania hamatiloba var. latistipula Hattori, 1975
  • Frullania hamatosetacea Grolle In Grolle & Meister, 2004
  • Frullania hamiflora Herzog & Clark In Clark & Schultz, 1953
  • Frullania handelii Verdoorn In Handel-Mazzetti, 1930
  • Frullania handel-Mazzettii Hattori, 1981
  • Frullania hariotana Stephani, 1911
  • Frullania harpantha Herzog, 1942
  • Frullania hasskarliana Lindenberg In G., L. & N., 1845
  • Frullania hasskarliana var. gracilis Hattori, 1986
  • Frullania hasskarliana var. integribractea (Verdoorn) Hattori, 1975
  • Frullania hasskarliana var. parvidentata Hattori, 1986
  • Frullania hattoriana Godfrey & Godfrey, 1980
  • Frullania hattoriantha Udar & Nath, 1981
  • Frullania hattorii Konrat & Braggins, 2003
  • Frullania hawaiiensis Miller, 1953
  • Frullania hebridensis Stephani, 1911
  • Frullania hedrantha Hattori & Kamimura, 1973
  • Frullania helleri Stephani, 1910
  • Frullania herzogiana Stephani, 1911
  • Frullania herzogii Hattori, 1955
  • Frullania heteromorpha Schiffner, 1889
  • Frullania hicksiae Hattori, 1984
  • Frullania hicksiae F. Litoralis Hattori, 1988
  • Frullania higuchii Yuzawa & Hattori In Yuzawa & Koike, 1994
  • Frullania himalayensis Stephani, 1910
  • Frullania hinoi Kamimura, 1982
  • Frullania hiroshii Hattori, 1980
  • Frullania hirtiflora Spruce, 1884
  • Frullania holostipula Hattori & Griffin Iii, 1978
  • Frullania hottana Hattori, 1976
  • Frullania howeana Stephani In Stephani & Watts, 1914
  • Frullania huerlimannii Hattori, 1976
  • Frullania huerlimannii var. dioica Hattori, 1984
  • Frullania humbertii Vanden Berghen, 1976
  • Frullania humilis Spruce, 1890
  • Frullania hypoleuca Nees In G., L. & N., 1843
  • Frullania hypoleucula Hattori, 1984

I

  • Frullania imerinensis Stephani, 1911
  • Frullania immersa Stephani, 1896
  • Frullania incisoduthiana Hattori In Mizutani, 1979
  • Frullania incisoduthiana var. parva Hattori In Mizutani, 1979
  • Frullania incisostipula Stephani, 1924
  • Frullania inconstans Verdoorn, 1930
  • Frullania inconstans F. Integrior Verdoorn, 1930
  • Frullania inconstans var. grossedentata Kamimura & Hattori, 1973
  • Frullania incumbens Mitten In Hooker, 1855
  • Frullania incurva Hattori, 1988
  • Frullania inflata Gottsche In G., L. & N., 1845
  • Frullania inflata var. communis Schuster, 1985
  • Frullania inflata var. dioica Hattori & Thaithong, 1978
  • Frullania inflexa Mitten, 1861
  • Frullania inflexiloba Hattori, 1984
  • Frullania inouei Hattori, 1980
  • Frullania integristipula (Nees) Nees In G., L. & N., 1845
  • Frullania integristipula var. emarginata Verdoorn, 1929
  • Frullania intermedia Subsp. (Intermedia 1980) F. Billardieriana (Nees & Montagne In Montagne) Verdoorn, 1930
  • Frullania intermedia Subsp. (Intermedia 1980) var. non-Apiculata Hattori, 1975
  • Frullania intermedia Subsp. (Intermedia) (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
  • Frullania intermedia Subsp. Morokensis (Stephani) Hattori, 1980
  • Frullania intermixta Colenso, 1889
  • Frullania intumescens (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
  • Frullania involuta Hampe Ex Stephani, 1911
  • Frullania involvens Hattori & Kamimura, 1973
  • Frullania iriomotensis Hattori, 1980
  • Frullania irregularis Hattori & Piippo, 1986
  • Frullania iwatsukii Hattori, 1972

J

  • Frullania jackii Subsp. (Jackii 1959) Gottsche In Gottsche & Rabenhorst, 1863
  • Frullania jackii Subsp. (Jackii 1959) F. Depauperata Grolle, 1970
  • Frullania jacobsii Hattori, 1986
  • Frullania jacquinotii Gottsche Ex Stephani, 1910
  • Frullania jelskii Loitlesberger, 1894
  • Frullania johnsonii Stephani, 1894
  • Frullania junghuhniana Gottsche In G., L. & N., 1845
  • Frullania junghuhniana var. bisexualis Hattori, 1976
  • Frullania junghuhniana var. tenella (Sande Lacoste) Grolle & Hattori In Hattori, 1982
  • Frullania junghuhniana var. tenella F. Monoica (Hattori) Hattori, 1986

K

  • Frullania kagoshimensis Subsp. (Kagoshimensis 1985) Stephani, 1910
  • Frullania kagoshimensis Subsp. (Kagoshimensis 1985) var. minor Kamimura, 1961
  • Frullania kagoshimensis Subsp. Hunanensis (Hattori) Hattori, 1985
  • Frullania kalimantanensis Hattori, 1986
  • Frullania kalimantanensis Piippo & Hattori In Piippo & Tan, 1992
  • Frullania kashyapii Verdoorn, 1932
  • Frullania kitagawana Hattori, 1984
  • Frullania klotzschii Nees Ex Stephani, 1911
  • Frullania koponenii Hattori, 1978
  • Frullania kunzei (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
  • Frullania kunzei var. maritima Schuster, 1991

L

  • Frullania laetevirens Hampe In G., L. & N., 1845
  • Frullania laevi-Periantha Bai & Gao, 2000
  • Frullania lancistyla Stephani, 1910
  • Frullania larjiana Singh & Singh, 2005
  • Frullania laticaulis Spruce, 1890
  • Frullania latiflora Spruce, 1884
  • Frullania latogaleata Herzog, 1948
  • Frullania laxiflora Spruce, 1884
  • Frullania laxiflora var. crossii Spruce, 1884
  • Frullania leana Austin, 1869
  • Frullania leeuwenii Verdoorn, 1930
  • Frullania lepida Hattori & Piippo, 1986
  • Frullania letestui Vanden Berghen, 1976
  • Frullania levieri Stephani, 1910
  • Frullania libera Nees In G., L. & N., 1845
  • Frullania lindbergiana Gottsche, 1863
  • Frullania lindenbergii Gottsche Ex Lehmann, 1844
  • Frullania lindenbergii var. (Beta ?) Fusca Gottsche In G., L. & N., 1847
  • Frullania lindeniana Stephani, 1911
  • Frullania lindmanii Stephani, 1897
  • Frullania linii Hattori, 1981
  • Frullania lobato-Hastata Stephani, 1911
  • Frullania lobulata (Hooker) Hooker & Nees In G., L. & N., 1845
  • Frullania longipinna Stephani, 1910
  • Frullania longistipula Stephani In Renauld & Cardot, 1891
  • Frullania longistipula var. apiculata Demaret & Vanden Berghen, 1950
  • Frullania longistyla Yuzawa & Hattori, 1988
  • Frullania loricata Pearson, 1891
  • Frullania loricata var. laxa Pearson, 1891
  • Frullania ludoviciae Stephani, 1908
  • Frullania lushanensis Hattori & Lin, 1985

M

  • Frullania macgregorii Stephani, 1894
  • Frullania macgregorii var. rostellula (Hattori) Hattori, 1982
  • Frullania macrocephala (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
  • Frullania macrophylla Hattori, 1980
  • Frullania macularis Taylor, 1846
  • Frullania madagascariensis Gottsche, 1882
  • Frullania madens Stephani, 1924
  • Frullania madothecoides Spruce, 1884
  • Frullania magellanica (Sprengel) Weber & Nees In G., L. & N., 1845
  • Frullania magellanica var. diminutiva Herzog, 1938
  • Frullania mammilligera Grolle, 2003
  • Frullania mammillosa Hattori, 1977
  • Frullania matafaoica Miller, 1981
  • Frullania mathanii Stephani, 1911
  • Frullania mauritiana Austin, 1869
  • Frullania maymyoensis Svihla, 1958
  • Frullania mcveanii Hattori, 1973
  • Frullania media (Hodgson) Hattori, 1983
  • Frullania megalostipa Spruce, 1884
  • Frullania meijeri Hattori, 1974
  • Frullania meridana Stephani, 1911
  • Frullania meyeniana Lindenberg In G., L. & N., 1845
  • Frullania meyeniana var. dioica Hattori, 1977
  • Frullania microauriculata Verdoorn, 1929
  • Frullania microauriculata var. rotundior Verdoorn, 1929
  • Frullania microcaulis Gola, 1923
  • Frullania microcephala Gottsche, 1863
  • Frullania microphylla (Gottsche In Gottsche & Rabenhorst) Pearson, 1894
  • Frullania microphylla F. Corticicola Schiffner, 1936
  • Frullania microphylla var. deciduifolia Grolle, 1970
  • Frullania microrhyncha Clark & Svihla, 1950
  • Frullania microscopica Pearson, 1922
  • Frullania minor Subsp. (Minor 1975) var. integribracteola Hattori, 1975
  • Frullania minor Subsp. Recurviloba Hattori, 1975
  • Frullania mirabilis Jack & Stephani In Stephani, 1892
  • Frullania miradorensis Lindenberg & Gottsche In G., L. & N., 1847
  • Frullania mizutanii Kamimura & Hattori, 1973
  • Frullania moniliata Subsp. Breviramea (Stephani) Verdoorn, 1930
  • Frullania moniliata Subsp. Moniliata Crandall-Stotler , Geissler & Stotler, 1987
  • Frullania moniliata Subsp. Moniliata F. Minshanensis (Hattori) Crandall-Stotler Et Alii, 1987
  • Frullania moniliata Subsp. Moniliata var. elongatistipula (Verdoorn) Crandall-Stotler Et Alii, 1987
  • Frullania moniliata Subsp. Moniliata var. vietnamica (Hattori) Crandall-Stotler Et Alii, 1987
  • Frullania monocera (Hooker & Taylor) Taylor In G., L. & N., 1845
  • Frullania monocera var. depauperata Hattori, 1984
  • Frullania monocera var. schiffneri (Verdoorn) Hattori, 1979
  • Frullania monoica Stephani, 1900
  • Frullania montagnei Gottsche In G., L. & N., 1845
  • Frullania montana Stephani, 1910
  • Frullania moritziana Lindenberg & Gottsche In G., L. & N, 1847
  • Frullania moritziana var. (Beta) Mexicana Gottsche, 1863
  • Frullania morobensis Hattori & Streimann, 1985
  • Frullania motoyana Stephani, 1911
  • Frullania mucronata (Lehmann & Lindenberg In Lehmann) Lehmann & Linde,nberg
  • Frullania mucronata var. submutica Lehmann & Lindenberg In G., L. & N. 1845 In G., L. & N., 1845
  • Frullania multilacera Subsp. (Multilacera 1987) Stephani, 1911
  • Frullania multilacera Subsp. (Multilacera 1987) var. lacerissima Hattori, 1975
  • Frullania multilacera Subsp. Gracilior Hattori, 1987
  • Frullania multilaceroides Hattori, 1987
  • Frullania muscicola Stephani, 1894
  • Frullania mutilata Stephani, 1911

N

  • Frullania nadeaudii Stephani, 1911
  • Frullania neocaledonica Engel In Engel & Merrill, 1999
  • Frullania neosheana Hattori, 1979
  • Frullania nepalensis (Sprengel) Lehmann & Lindenberg In G., L. & N., 1845
  • Frullania neurota Taylor, 1846
  • Frullania nicholsonii Hodgson, 1949
  • Frullania nigricaulis (R., B. & N.) Nees In G., L. & N., 1845
  • Frullania nigricaulis var. elongata Verdoorn, 1930
  • Frullania nisquallensis Sullivant, 1849
  • Frullania nivimontana Hattori, 1982
  • Frullania nobilis Stephani, 1894
  • Frullania nobilis var. cochleata (Stephani) Hattori In Hattori & Streimann, 1985
  • Frullania nodulosa (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
  • Frullania notarsii Stephani, 1911
  • Frullania novocurvirostris Hattori, 1981
  • Frullania novoguineensis Schiffner, 1890

O

  • Frullania oahuensis Hampe In G., L. & N., 1843
  • Frullania oakesiana Subsp. (Oakesiana 1992) Austin, 1870
  • Frullania oakesiana Subsp. Takayuensis (Stephani) Schuster, 1992
  • Frullania obovata Hattori, 1982
  • Frullania obscurifolia Mitten, 1879
  • Frullania ocanniensis Stephani, 1924
  • Frullania ocellata Kamimura & Hattori, 1973
  • Frullania odontostipa Spruce, 1890
  • Frullania okinawensis Kamimura, 1982
  • Frullania onraedtii Vanden Berghen, 1976
  • Frullania orbicularis Austin, 1869
  • Frullania orientalis Sande Lacoste, 1855
  • Frullania orinocensis Spruce, 1883
  • Frullania ornithocephala (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
  • Frullania ornithocephala F. Magnilobula Hattori, 1976
  • Frullania ornithocephala F. Retusa Hattori, 1976
  • Frullania ornithocephala F. Teres (Sande Lacoste) Verdoorn, 1929
  • Frullania ornithocephala var. maior Nees, 1845
  • Frullania ornithocephala var. minor Nees, 1845
  • Frullania ornithocephala var. pilosa Verdoorn, 1929
  • Frullania ornithocephala var. tuberculosa Hattori, 1974
  • Frullania osculatiana Notaris, 1855
  • Frullania osumiensis (Hattori) Hattori In Iwatsuki & Hattori, 1956
  • Frullania osumiensis var. dentata Kamimura, 1970

P

  • Frullania pachyderma Hattori, 1978
  • Frullania pallide-Virens Stephani, 1911
  • Frullania pallidula Hattori, 1988
  • Frullania pancheri Gottsche Ex Stephani, 1894
  • Frullania papillata Stephani, 1911
  • Frullania papillilobula Hattori, 1975
  • Frullania papuana Verdoorn, 1930
  • Frullania papulosa Stephani, 1911
  • Frullania paradoxa Lehmann & Lindenberg In G., L. & N., 1845
  • Frullania paradoxa F. Cornistipula Herzog, 1952
  • Frullania parhami (Schuster) Schuster, 1992
  • Frullania pariharii Hattori & Thaithong, 1978
  • Frullania pariharii F. Intermedia Hattori & Thaithong, 1978
  • Frullania parvifolia Stephani, 1910
  • Frullania parvistipula Stephani, 1910
  • Frullania patagonica Stephani, 1910
  • Frullania patula Mitten In Hooker, 1855
  • Frullania pauciramea Stephani, 1911
  • Frullania pauciramea var. pauciramella Hattori & Piippo, 1986
  • Frullania paucirameoides Hattori & Piippo, 1986
  • Frullania pearceana Stephani, 1911
  • Frullania pedicellata Stephani, 1897
  • Frullania pentapleura Subsp. (Pentapleura 1988) Taylor, 1846
  • Frullania peruviana Gottsche In G., L. & N., 1846
  • Frullania phalangiflora Stephani, 1916
  • Frullania philippinensis Stephani, 1911
  • Frullania physantha Mitten Mitten, 1861
  • Frullania pilibracteola Hattori, 1977
  • Frullania piliflora Stephani, 1911
  • Frullania piliflora var. appendiculata Herzog, 1951
  • Frullania pilistipula Stephani, 1911
  • Frullania piptophylla Hattori, 1980
  • Frullania piptophylla var. minor Hattori, 1986
  • Frullania piptophylloides Hattori, 1980
  • Frullania pittier I Stephani, 1892
  • Frullania plana Sullivant, 1849
  • Frullania planifolia Stephani, 1910
  • Frullania platycalyx Herzog, 1952
  • Frullania platyphylla Colenso, 1889
  • Frullania pluricarinata Gottsche, 1864
  • Frullania pocsantha Thaithong & Hattori, 1977
  • Frullania polyclada Colenso, 1889
  • Frullania polyptera Taylor, 1846
  • Frullania polyptera var. angustata (Mitten) Hattori, 1975
  • Frullania polysticta Lindenberg In G., L. & N., 1845
  • Frullania ponapensis Hattori & Koike In Koike, 1994
  • Frullania pringlei Fulford & Sharp, 1990
  • Frullania probosciphora Taylor, 1846
  • Frullania prominula Hattori & Streimann, 1985
  • Frullania propaginea Hattori & Streimann, 1985
  • Frullania pseudericoides Hattori, 1982
  • Frullania pseudericoides Hattori, 1986
  • Frullania pseudomeyeniana Hattori, 1986
  • Frullania pseudomonocera Hattori, 1986
  • Frullania pseudoschensiana Hattori, 1980
  • Frullania pseudoschensiana var. darjeelingensis Hattori, 1981
  • Frullania ptychantha Montagne, 1843
  • Frullania pulchella Herzog, 1954
  • Frullania pullei Verdoorn, 1930
  • Frullania pulogensis Stephani, 1911
  • Frullania punctata Reimers, 1931
  • Frullania purpurea Stephani, 1911
  • Frullania pusilla Mitten, 1871
  • Frullania pycnantha (Hooker & Taylor) Taylor In G., L. & N., 1845
  • Frullania pycnoclada Grolle In Grolle & Meister, 2004
  • Frullania pyricalycina Stephani, 1910

Q

  • Frullania queenslandica Stephani, 1910
  • Frullania quillotensis (Nees & Montagne) Nees & Montagne In Montagne In Alcide D'orbigny, 1839
  • Frullania quillotensis F. Flaccida Herzog, 1954
  • Frullania quillotensis F. Leiantha Herzog, 1954

R

  • Frullania rabenhorstii Stephani, 1911
  • Frullania ramuligera (Nees) Montagne, 1842
  • Frullania recurvistipula Hattori, 1975
  • Frullania reflexistipula Sande Lacoste, 1853
  • Frullania reflexistipula var. novocaledonica Hattori, 1977
  • Frullania reflexistipula var. squarrosa Hattori & Piippo, 1986
  • Frullania regularis Schiffner, 1890
  • Frullania reicheana Stephani, 1910
  • Frullania reimersii Verdoorn, 1930
  • Frullania remotidens Hattori, 1872
  • Frullania remotifolia Stephani, 1911
  • Frullania remotiloba Stephani, 1894
  • Frullania repanda Gottsche, 1864
  • Frullania repandistipula Subsp. (Repandistipula 1975) Sande Lacoste, 1853
  • Frullania repandistipula Subsp. Queenslandica Hattori, 1987
  • Frullania repandistipula Subsp. Spinibractea Hattori, 1975
  • Frullania reptans var. integristipula Nicholson, 1925
  • Frullania retusa Mitten, 1861
  • Frullania retusa var. gymnantha Hattori & Thaithong, 1978
  • Frullania retusa var. hirsuta Hattori & Thaithong, 1978
  • Frullania rhystocolea Herzog Ex Verdoorn In Handel-Mazzetti, 1930
  • Frullania rhytidantha Hattori, 1980
  • Frullania rigescens Spruce, 1884
  • Frullania rigida Stephani, 1910
  • Frullania ringens Spruce, 1884
  • Frullania riojaneirensis (Raddi) Ångström, 1876
  • Frullania riparia Hampe Ex Lehmann, 1838
  • Frullania rizalli Piippo & Hattori, 1992
  • Frullania rostellata Mitten In Hooker, 1867
  • Frullania rostrata (Hooker & Taylor) Hooker & Taylor In G., L. & N., 1845
  • Frullania rubella Gottsche Ex Stephani, 1889
  • Frullania rubella var. elongata (Stephani) Hattori, 1983
  • Frullania rudolfiana Hattori, 1972
  • Frullania rupicola Stephani, 1924

S

  • Frullania sabahana Hattori, 1976
  • Frullania sabaliana Schuster, 1983
  • Frullania sabanetica Gottsche, 1864
  • Frullania sachapatensis Stephani, 1911
  • Frullania sackawana Stephani, 1897
  • Frullania sackawana var. minor Kamimura, 1982
  • Frullania saipanensis Hattori & Koike In Koike, 1994
  • Frullania sarawakensis Hattori, 1976
  • Frullania scalaris Hattori, 1977
  • Frullania scandens Montagne, 1843
  • Frullania schaefer-Verwimpii Yuzawa & Hattori, 1989
  • Frullania schensiana Massalongo, 1897
  • Frullania schiffneri Verdoorn, 1929
  • Frullania schimperi Nees In G., L. & N., 1845
  • Frullania schimperi var. laciniata Vanden Berghen, 1976
  • Frullania schumannii (Caspary) Grolle, 1981
  • Frullania schusteri Hattori, 1988
  • Frullania schusteriana Hattori, 1972
  • Frullania scottiana Hattori, 1987
  • Frullania sebastianopolitana var. (Delta) Gottsche, 1864
  • Frullania selwyniana Pearson, 1890
  • Frullania semienana Gola, 1914
  • Frullania semivillosa Lindenberg & Gottsche In G., L. & N., 1847
  • Frullania semivillosa var. (Beta) Glabra Lindenberg & Gottsche In G., L. & N., 1847
  • Frullania sergiae Sim-Sim, Fontihna, Mues & Lion, 2000
  • Frullania seriata Gottsche Ex Stephani, 1889
  • Frullania seriatifolia Stephani, 1894
  • Frullania serrata Subsp. (Serrata 1982) Gottsche In G., L. & N., 1845
  • Frullania serrata Subsp. (Serrata 1982) F. Crispulodentata Verdoorn, 1929
  • Frullania serrata Subsp. (Serrata 1982) F. Vogelkopensis Hattori, 1982
  • Frullania serrata Subsp. (Serrata 1982) var. ceramensis Hattori, 1986
  • Frullania serrata Subsp. (Serrata 1982) var. hamatispina (Hattori) Hattori, 1982
  • Frullania serrata Subsp. (Serrata 1982) var. oceanica Verdoorn, 1937
  • Frullania serrata Subsp. (Serrata 1982) var. pertenuis (Nees) Gottsche In G., L. & N., 1845
  • Frullania serrata Subsp. Grolleana Grolle & Piippo, 1984
  • Frullania serrata Subsp. Spinistipula (Hattori) Hattori, 1982
  • Frullania setacea Hattori, 1988
  • Frullania setigera Stephani, 1894
  • Frullania shanensis Svihla, 1957
  • Frullania sharpantha Udar & Kumar, 1983
  • Frullania sharpii Hattori, 1974
  • Frullania sheana Hattori, 1979
  • Frullania simmondsii Stephani In Stephani & Watts, 1914
  • Frullania sinensis Stephani In Levier, 1906
  • Frullania sinosphaerantha Hattori & Lin, 1985
  • Frullania sinuata Subsp. Sinuata 1975 Sande Lacoste, 1853
  • Frullania sinuata Subsp. Sinuata F. Parvistylata Hattori In Grolle & Piippo, 1984
  • Frullania socotrana Mitten In Balfour, 1888
  • Frullania solanderiana Colenso, 1889
  • Frullania spathulistipa Stephani, 1910
  • Frullania speciosa Herzog, 1949
  • Frullania spegazzinii Reiner, 1988
  • Frullania sphaerantha Hattori, 1980
  • Frullania sphaerocephala Spruce, 1884
  • Frullania sphaerolobulata Lin In Lin & Chen, 1997
  • Frullania spicata (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
  • Frullania spinifera Taylor, 1846
  • Frullania spinigastria Hattori, 1979
  • Frullania spiniplica Hattori, 1972
  • Frullania spinistipula Stephani, 1911
  • Frullania spongiosa Stephani, 1894
  • Frullania squamuligera Spruce, 1884
  • Frullania squarrosula (Hooker & Taylor) Taylor In G., L. & N., 1845
  • Frullania standaerti Stephani, 1910
  • Frullania steereana Hattori, 1987
  • Frullania stenostipa Spruce, 1884
  • Frullania streimannii Hattori, 1983
  • Frullania stylifera (Schuster) Schuster, 1992
  • Frullania subcaduca Hattori, 1974
  • Frullania subclavata Stephani, 1910
  • Frullania subdentata Stephani, 1911
  • Frullania subdentata F. Longistipula Hattori, 1978
  • Frullania subdilatata Massalongo In Levier, 1906
  • Frullania subhampeana Hodgson, 1949
  • Frullania subincumbens Hattori, 1987
  • Frullania sublignosa Stephani, 1894
  • Frullania submultilacera Hattori & Koike In Koike, 1994
  • Frullania subnigricaulis Hattori, 1973
  • Frullania subnigricaulis var. subtruncata Hattori, 1975
  • Frullania subocellata Hattori, 1986
  • Frullania subpedicellata Hattori, 1980
  • Frullania subpilibracteola Hattori, 1977
  • Frullania subpyricalycina Herzog, 1954
  • Frullania subrostrata Hattori, 1972
  • Frullania subsquarrosa Hattori, 1972
  • Frullania subtilissima Lindenberg In G., L. & N., 1845
  • Frullania subtropica Stephani, 1910
  • Frullania subtruncata Stephani, 1910
  • Frullania subvalida Hattori & Thaithong, 1978
  • Frullania supradecomposita (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
  • Frullania svihlana Hattori, 1983

T

  • Frullania tagawana (Hattori, Thaithong & Kitawana) Hattori In Hattori & Lin, 1985
  • Frullania taiheizana Horikawa, 1934
  • Frullania tamarisci (L.) Dumortier, 1835
  • Frullania tamarisci var. atlantica Schiffner, 1936
  • Frullania tamarisci var. atrovirens Carrington, 1863
  • Frullania tamarisci var. commutata Nees In G., L. & N., 1845
  • Frullania tamarisci var. cornubica Carrington In Carrington & Pearson, 1878
  • Frullania tamarisci var. laxa Nees In G., L. & N., 1845
  • Frullania tamarisci var. mediterranea (De Notaris) Nees In G., L. & N., 1845
  • Frullania tamarisci var. robusta Lindberg, 1875
  • Frullania tamarisci var. sardoa (De Notaris) De Notaris, 1865
  • Frullania tamarisci var. schiffneri Nicholson, 1933
  • Frullania tamsuina Stephani, 1910
  • Frullania taradakenis Stephani, 1910
  • Frullania taxodiocola Schuster, 1983
  • Frullania teneriffae (Weber) Nees, 1838
  • Frullania teneriffae F. Explanata (Macvicar) Bisang, Schumacker, Sérgio & Grolle, 1988
  • Frullania teneriffae var. germana (Taylor) Bisang, Schumacker, Sérgio & Grolle, 1988
  • Frullania tenuirostris Stephani, 1911
  • Frullania ternatensis Gottsche In G., L. & N., 1846
  • Frullania ternatensis var. non-Appendiculata Hattori, 1974
  • Frullania tetraptera Nees & Montagne In Montagne, 1838
  • Frullania thiersiae Hattori, 1988
  • Frullania tixieri Hattori, 1976
  • Frullania tjibodensis Hattori & Thaithong, 1977
  • Frullania togashiana Hattori, 1975
  • Frullania tongariroense Colenso, 1889
  • Frullania trianae Gottsche, 1864
  • Frullania tricarinata Sande Lacoste, 1856
  • Frullania trichodes Mitten, 1862
  • Frullania trigona Clark, Jovet-Ast & Frye, 1947
  • Frullania trinervis (Lehmann & Lindenberg) Drège, 1843
  • Frullania triquetra Lindenberg & Gottsche In G., L. & N., 1847
  • Frullania trollii Herzog, 1942
  • Frullania truncata Caspary, 1887
  • Frullania tubercularis Hattori & Lin, 1985
  • Frullania tunguraguana Clark & Frye, 1952
  • Frullania turfosa Lindenberg & Gottsche In G., L. & N., 1847
  • Frullania tuyamae Hattori & Thaithong, 1978

U

  • Frullania udarii Nath & Singh, 2006
  • Frullania umbonata Mitten Ex Stephani, 1911
  • Frullania undulata Kamimura, 1961
  • Frullania usambarana Schiffner Ex Stephani, 1894
  • Frullania usambarana var. reducta Vanden Berghen, 1976
  • Frullania usamiensis Stephani, 1897
  • Frullania utriculata Stephani, 1894

V

  • Frullania vaga Mitten In Seemann, 1871
  • Frullania vaginata (Swartz) Nees In G., L. & N., 1846
  • Frullania valdiviensis Jack & Stephani In Stephani, 1894
  • Frullania valida Stephani, 1910
  • Frullania valparaisiana Lehmann, 1857
  • Frullania vandenberghenii Pocs In Bizot & Pocs, 1979
  • Frullania van-Zantenii Kamimura & Hattori, 1973
  • Frullania variabilis Stephani, 1910
  • Frullania varians Caspary, 1887
  • Frullania variegata Stephani, 1894
  • Frullania venusta Hattori, 1974
  • Frullania verdoorniana Hattori, 1973
  • Frullania victoriensis Stephani, 1910
  • Frullania vitalii Yuzawa & Hattori, 1988
  • Frullania vittata Hattori, 1974
  • Frullania vittata F. Denticulata Hattori, 1980
  • Frullania vittiana Hattori, 1987

W

  • Frullania wagneri Stephani, 1910
  • Frullania wairua Konrat & Braggins, 2005
  • Frullania wangii Hattori & Lin, 1985
  • Frullania warnckeana Hattori, 1974
  • Frullania warnckeana var. dentosa Hattori, 1977
  • Frullania weberbaueri Stephani, 1911
  • Frullania winteri Stephani, 1910
  • Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa, 1991

Y

  • Frullania yorkiana Stephani, 1911
  • Frullania yuennanensis Stephani, 1894
  • Frullania yuennanensis var. siamensis (Hatt. et al.) Hattori & Lin, 1985
  • Frullania yusawae Kamimura, 1970
  • Frullania yuzawana Hattori, 1981

Z

  • Frullania zangii Hattori & Lin, 1985
  • Frullania zennoskeana Hattori, 1984

Extinct species

  • Frullania baerlocheri Heinrichs et al., 2012 Burmese amber, Myanmar, Cenomanian
  • Frullania cretacea Hentschel et al., 2009 Burmese amber, Myanmar, Cenomanian
  • Frullania partita Li et al, 2020 Burmese amber, Myanmar, Cenomanian
  • Frullania pinnata Heinrichs et al., 2017 Burmese amber, Myanmar, Cenomanian

References

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