Khodzhakul Formation

The Khodzhakul Formation is a Cenomanian aged geologic formation in Uzbekistan. Dinosaur remains are among the fossils that have been recovered from the formation.[1] As well as those of Mammals.[2] It is part of the same stratigraphic succession as the overlying Bissekty Formation.

Khodzhakul Formation
Stratigraphic range: Cenomanian
~93–89 Ma
↓
TypeGeological formation
UnderliesTamdytruba & Bissekty Formations
OverliesBortesken Formation
Lithology
PrimarySandstone
Location
Coordinates40.1°N 65.8°E / 40.1; 65.8
Approximate paleocoordinates35.9°S 65.5°E / -35.9; 65.5
RegionKarakalpakstan
Country Uzbekistan
Khodzhakul Formation (Uzbekistan)

Fossil content

One of the largest mammals of the Mesozoic, Oxlestes, occurred in this formation, and may have predated on dinosaur species.[3]

Dinosaurs
Genus Species Location Stratigraphic position Material Notes Images

Asiaceratops[4]

A. salsopaludalis[4]

"Teeth, cranial fragments, phalanx."[5]

Gilmoreosaurus[6]

G. atavus[6]

Horezmavis[7]

H. eocretaceous[7]

"Tarsometatarsus."[8]

Kulceratops[4]

K. kulensis[4]

"Fragmentary material."[5]

Therizinosauroidea

Indeterminate[9]

Upper

Pectinodon[4]

P. Sp[4]
Mammals[2]
Genus Species Location Stratigraphic position Material Notes Images
Bobolestes B. zenge Maxilla and dentary fragments Zalambdalestoidea
Eozhelestes E. mangit dentary fragments and teeth
Sheikhdzheilia S. rezvyii Maxilla and dentary fragments ?Zhelestidae
Zalambdalestidae Indeterminate Right petrosal
‘‘Zhelestidae’’ Indeterminate Left dentary and dentary fragment Large taxon
Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
 Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

See also
  • List of dinosaur-bearing rock formations

References
  1. Weishampel, et al., 2004, "Dinosaur distribution." pp.517-607
  2. Averianov, Alexander; Archibald, J. David (August 2005). "Mammals from the mid-Cretaceous Khodzhakul Formation, Kyzylkum Desert, Uzbekistan". Cretaceous Research. 26 (4): 593–608. doi:10.1016/j.cretres.2005.03.007.
  3. Michael J. Benton,Mikhail A. Shishkin, David M. Unwin, The Age of Dinosaurs in Russia and Mongolia, Cambridge University Press, 04/12/2003 - 740 páginas
  4. "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Upper Khodzhakul Formation," in Weishampel, et al. (2004). Page 594.
  5. "Table 22.1," in Weishampel, et al. (2004). Page 480.
  6. Listed as "? Gilmoreosaurus sp." in "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Upper Khodzhakul Formation," in Weishampel, et al. (2004). Page 594.
  7. "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Lower Khodzhakul Formation," in Weishampel, et al. (2004). Page 563.
  8. "Table 11.1," in Weishampel, et al. (2004). Page 214.
  9. Sues, Hans-Dieter; Averianov, Alexander (2016). "Therizinosauroidea (Dinosauria: Theropoda) from the Upper Cretaceous of Uzbekistan". Cretaceous Research. 59: 155–178. doi:10.1016/j.cretres.2015.11.003.

Bibliography
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