List of birds of Telangana

The state of Telangana, India, has 492 bird species within its boundaries.

The official state bird of Telangana is the pala pitta,[1] also known as the Indian roller, Coracias benghalensis.

This list's taxonomic treatment (designation and sequence of orders, families and species) and nomenclature (common and scientific names) follow the conventions of the IOC World Bird List, version 10.2. This list also uses British English throughout. Any bird names or other wording follows that convention.

The following tags have been used to highlight several categories. The commonly occurring native species do not fit within any of these categories.

  • (A) Accidental - Also known as a rarity, it refers to a species that rarely or accidentally occurs in Telangana-typically less than ten confirmed records.


Ducks, geese, and swans
Bar-headed goose
Northern shoveller

Order: Anseriformes   Family: Anatidae

Anatidae includes the ducks and most duck-like waterfowl, such as geese and swans. These birds are adapted to an aquatic existence with webbed feet, flattened bills, and feathers that are excellent at shedding water due to an oily coating.

Pheasants and allies
Grey francolin

Order: Galliformes   Family: Phasianidae

The Phasianidae are a family of terrestrial birds which consists of quails, partridges, snowcocks, francolins, spurfowls, tragopans, monals, pheasants, peafowls and jungle fowls. In general, they are plump (although they vary in size) and have broad, relatively short wings.

Nightjars
Indian nightjar

Order: Caprimulgiformes   Family: Caprimulgidae

Nightjars are medium-sized nocturnal birds that usually nest on the ground. They have long wings, short legs and very short bills. Most have small feet, of little use for walking, and long pointed wings. Their soft plumage is camouflaged to resemble bark or leaves.

Treeswifts
Crested treeswift

Order: Apodiformes   Family: Hemiprocnidae

The treeswifts, also called crested swifts, are closely related to the true swifts. They differ from the other swifts in that they have crests, long forked tails and softer plumage.

Swifts

Order: Apodiformes   Family: Apodidae

Swifts are small birds which spend the majority of their lives flying. These birds have very short legs and never settle voluntarily on the ground, perching instead only on vertical surfaces. Many swifts have long swept-back wings which resemble a crescent or boomerang.

Bustards
Great Indian bustard

Order: Otidiformes   Family: Otididae

Bustards are large terrestrial birds mainly associated with dry open country and steppes in the Old World. They are omnivorous and nest on the ground. They walk steadily on strong legs and big toes, pecking for food as they go. They have long broad wings with "fingered" wingtips and striking patterns in flight. Many have interesting mating displays.

Cuckoos
Jacobin cuckoo
Greater coucal

Order: Cuculiformes   Family: Cuculidae

The family Cuculidae includes cuckoos, roadrunners and anis. These birds are of variable size with slender bodies, long tails and strong legs. Many are brood parasites.

Sandgrouse

Order: Pterocliformes   Family: Pteroclidae

Sandgrouse have small, pigeon like heads and necks, but sturdy compact bodies. They have long pointed wings and sometimes tails and a fast direct flight. Flocks fly to watering holes at dawn and dusk. Their legs are feathered down to the toes.

Pigeons and doves
Laughing dove
Painted sandgrouse

Order: Columbiformes   Family: Columbidae

Pigeons and doves are stout-bodied birds with short necks and short slender bills with a fleshy cere.

Rails, crakes, and coots
White-breasted waterhen

Order: Gruiformes   Family: Rallidae

Rallidae is a large family of small to medium-sized birds which includes the rails, crakes, coots and gallinules. Typically they inhabit dense vegetation in damp environments near lakes, swamps or rivers. In general they are shy and secretive birds, making them difficult to observe. Most species have strong legs and long toes which are well adapted to soft uneven surfaces. They tend to have short, rounded wings and to be weak fliers.

Cranes
Demoiselle crane

Order: Gruiformes   Family: Gruidae

Cranes are large, long-legged and long-necked birds. Unlike the similar-looking but unrelated herons, cranes fly with necks outstretched, not pulled back. Most have elaborate and noisy courting displays or "dances".

Grebes
Little grebe

Order: Podicipediformes   Family: Podicipedidae

Grebes are small to medium-large freshwater diving birds. They have lobed toes and are excellent swimmers and divers. However, they have their feet placed far back on the body, making them quite ungainly on land.

Flamingos
Greater flamingo

Order: Phoenicopteriformes   Family: Phoenicopteridae

Flamingos are gregarious wading birds, usually 3 to 5 feet (0.9 to 1.5 m) tall, found in both the Western and Eastern Hemispheres. Flamingos filter-feed on shellfish and algae. Their oddly shaped beaks are specially adapted to separate mud and silt from the food they consume and, uniquely, are used upside-down.

Buttonquail

Order: Charadriiformes   Family: Turnicidae

The buttonquail are small, drab, running birds which resemble the true quails. The female is the brighter of the sexes and initiates courtship. The male incubates the eggs and tends the young.

Stone-curlews and thick-knees
Great stone-curlew

Order: Charadriiformes   Family: Burhinidae

The stone-curlews are a group of largely tropical waders in the family Burhinidae. They are found worldwide within the tropical zone, with some species also breeding in temperate Europe and Australia. They are medium to large waders with strong black or yellow-black bills, large yellow eyes and cryptic plumage. Despite being classed as waders, most species have a preference for arid or semi-arid habitats.

Stilts and avocets
Black-winged stilt

Order: Charadriiformes   Family: Recurvirostridae

Recurvirostridae is a family of large wading birds, which includes the avocets and stilts. The avocets have long legs and long up-curved bills. The stilts have extremely long legs and long, thin, straight bills.

Plovers
Red-wattled lapwing
Yellow-wattled lapwing

Order: Charadriiformes   Family: Charadriidae

The family Charadriidae includes the plovers, dotterels and lapwings. They are small to medium-sized birds with compact bodies, short, thick necks and long, usually pointed, wings. They are found in open country worldwide, mostly in habitats near water.

Painted-snipes

Order: Charadriiformes   Family: Rostratulidae

Painted-snipes are short-legged, long-billed birds similar in shape to the true snipes, but more brightly coloured.

Jacanas

Order: Charadriiformes   Family: Jacanidae

The jacanas are a group of tropical waders in the family Jacanidae. They are found throughout the tropics. They are identifiable by their huge feet and claws which enable them to walk on floating vegetation in the shallow lakes that are their preferred habitat.

Sandpipers and snipes
Common sandpiper
Common snipe
Ruff

Order: Charadriiformes   Family: Scolopacidae

Scolopacidae is a large diverse family of small to medium-sized shorebirds including the sandpipers, curlews, godwits, shanks, tattlers, woodcocks, snipes, dowitchers and phalaropes. The majority of these species eat small invertebrates picked out of the mud or soil. Variation in length of legs and bills enables multiple species to feed in the same habitat, particularly on the coast, without direct competition for food.

Coursers and pratincoles

Order: Charadriiformes   Family: Glareolidae

Glareolidae is a family of wading birds comprising the pratincoles, which have short legs, long, pointed wings and long, forked tails, and the coursers, which have long legs, short wings and long, pointed bills which curve downwards.

Gulls, terns, and skimmers
Brown-headed gull
River tern
Little tern

Order: Charadriiformes   Family: Laridae

Laridae is a family of medium to large seabirds, the gulls, terns, and skimmers. Gulls are typically grey or white, often with black markings on the head or wings. They have stout, longish bills and webbed feet. Terns are a group of generally medium to large seabirds typically with grey or white plumage, often with black markings on the head. Most terns hunt fish by diving but some pick insects off the surface of fresh water. Terns are generally long-lived birds, with several species known to live in excess of 30 years.

Storks
Painted stork

Order: Ciconiiformes   Family: Ciconiidae

Storks are large, long-legged, long-necked, wading birds with long, stout bills. Storks are mute, but bill-clattering is an important mode of communication at the nest. Their nests can be large and may be reused for many years. Many species are migratory.

Anhingas and darters
Oriental darter

Order: Suliformes   Family: Anhingidae

Anhingas and darters are often called "snake-birds" because of their long thin neck, which gives a snake-like appearance when they swim with their bodies submerged. The males have black and dark-brown plumage, an erectile crest on the nape and a larger bill than the female. The females have much paler plumage especially on the neck and underparts. The darters have completely webbed feet and their legs are short and set far back on the body. Their plumage is somewhat permeable, like that of cormorants, and they spread their wings to dry after diving.

Cormorants and shags
Little cormorant

Order: Suliformes   Family: Phalacrocoracidae

Phalacrocoracidae is a family of medium to large coastal, fish-eating seabirds that includes cormorants and shags. Plumage colouration varies, with the majority having mainly dark plumage, some species being black-and-white and a few being colourful.

Ibises and spoonbills
Black-headed ibis

Order: Pelecaniformes   Family: Threskiornithidae

Threskiornithidae is a family of large terrestrial and wading birds which includes the ibises and spoonbills. They have long, broad wings with 11 primary and about 20 secondary feathers. They are strong fliers and despite their size and weight, very capable soarers.

Herons and bitterns
Black-crowned night heron
Indian pond heron

Order: Pelecaniformes   Family: Ardeidae

The family Ardeidae contains the bitterns, herons and egrets. Herons and egrets are medium to large wading birds with long necks and legs. Bitterns tend to be shorter-necked and more wary. Members of Ardeidae fly with their necks retracted, unlike other long-necked birds such as storks, ibises and spoonbills.

Pelicans
Spot-billed pelican

Order: Pelecaniformes   Family: Pelecanidae

Pelicans are large water birds with a distinctive pouch under their beak. As with other members of the order Pelecaniformes, they have webbed feet with four toes.

Ospreys
Western osprey

Order: Accipitriformes   Family: Pandionidae

The family Pandionidae contains usually only one species, the osprey. The osprey is a medium-large raptor which is a specialist fish-eater with most taxonomic authorities consider a worldwide distribution. A few authorities, including the IOC, split the bird into 2 species.

Kites, hawks, and eagles
Black-winged kite
Brahminy kite
Black kite

Order: Accipitriformes   Family: Accipitridae

Accipitridae is a family of birds of prey, which includes hawks, eagles, kites, harriers and Old World vultures. These birds have powerful hooked beaks for tearing flesh from their prey, strong legs, powerful talons and keen eyesight.

Barn owls

Order: Strigiformes   Family: Tytonidae

Barn owls are medium to large owls with large heads and characteristic heart-shaped faces. They have long strong legs with powerful talons.

Owls
Spotted owlet
Indian eagle-owl

Order: Strigiformes   Family: Strigidae

The typical owls are small to large solitary nocturnal birds of prey. They have large forward-facing eyes and ears, a hawk-like beak and a conspicuous circle of feathers around each eye called a facial disk.

Trogons

Order: Trogoniformes   Family: Trogonidae

The family Trogonidae includes trogons and quetzals. Found in tropical woodlands worldwide, they feed on insects and fruit, and their broad bills and weak legs reflect their diet and arboreal habits. Although their flight is fast, they are reluctant to fly any distance. Trogons have soft, often colourful, feathers with distinctive male and female plumage.

Hoopoes
Eurasian hoopoe

Order: Bucerotiformes   Family: Upupidae

Hoopoes have black, white and orangey-pink colouring with a large erectile crest on their head.

Hornbills
Indian grey hornbill

Order: Bucerotiformes   Family: Bucerotidae

Hornbills are a group of birds whose bill is shaped like a cow's horn, but without a twist, sometimes with a casque on the upper mandible. Frequently, the bill is brightly coloured.

Rollers

Order: Coraciiformes   Family: Coraciidae

Rollers resemble crows in size and build, but are more closely related to the kingfishers and bee-eaters. They share the colourful appearance of those groups with blues and browns predominating. The two inner front toes are connected, but the outer toe is not.

Kingfishers
Pied kingfishers

Order: Coraciiformes   Family: Alcedinidae

Kingfishers are medium-sized birds with large heads, long, pointed bills, short legs and stubby tails.

Bee-eaters
Green bee-eater

Order: Coraciiformes   Family: Meropidae

The bee-eaters are a group of near passerine birds in the family Meropidae. Most species are found in Africa but others occur in southern Europe, Madagascar, Australia and New Guinea. They are characterised by richly coloured plumage, slender bodies and usually elongated central tail feathers. All are colourful and have long downturned bills and pointed wings, which give them a swallow-like appearance when seen from afar.

Asian barbets

Order: Piciformes   Family: Megalaimidae

The barbets are plump birds, with short necks and large heads. They get their name from the bristles which fringe their heavy bills. Most species are brightly coloured.

Woodpeckers
White-bellied woodpecker

Order: Piciformes   Family: Picidae

Woodpeckers are small to medium-sized birds with chisel-like beaks, short legs, stiff tails and long tongues used for capturing insects. Some species have feet with two toes pointing forward and two backward, while several species have only three toes. Many woodpeckers have the habit of tapping noisily on tree trunks with their beaks.

Caracaras and falcons

Order: Falconiformes   Family: Falconidae

Falconidae is a family of diurnal birds of prey. They differ from hawks, eagles, and kites in that they kill with their beaks instead of their talons.

Old World parrots
Plum-headed parakeet

Order: Psittaciformes   Family: Psittaculidae

Characteristic features of parrots include a strong curved bill, an upright stance, strong legs, and clawed zygodactyl feet. Many parrots are vividly coloured, and some are multi-coloured. In size they range from 8 cm (3.1 in) to 1 m (3.3 ft) in length. Old World parrots are found from Africa east across south and southeast Asia and Oceania to Australia and New Zealand.

Pittas
Indian pitta

Order: Passeriformes   Family: Pittidae

Pittas are medium-sized by passerine standards and are stocky, with fairly long, strong legs, short tails and stout bills. Many are brightly coloured. They spend the majority of their time on wet forest floors, eating snails, insects and similar invertebrates.

Vangas, helmetshrikes, woodshrikes, and shrike-flycatchers

Order: Passeriformes   Family: Vangidae

The woodshrikes are similar in build to the shrikes.

Woodswallows, butcherbirds, and peltops

Order: Passeriformes   Family: Artamidae

The woodswallows are soft-plumaged, somber-coloured passerine birds. They are smooth, agile flyers with moderately large, semi-triangular wings.

Ioras

Order: Passeriformes   Family: Aegithinidae

The ioras are bulbul-like birds of open forest or thorn scrub, but whereas that group tends to be drab in colouration, ioras are sexually dimorphic, with the males being brightly plumaged in yellows and greens.

Cuckooshrikes
Small minivet

Order: Passeriformes   Family: Campephagidae

The cuckooshrikes are small to medium-sized passerine birds. They are predominantly greyish with white and black, although some species are brightly coloured.

Shrikes

Order: Passeriformes   Family: Laniidae Shrikes are passerine birds known for their habit of catching other birds and small animals and impaling the uneaten portions of their bodies on thorns. A typical shrike's beak is hooked, like a bird of prey.

Figbirds, orioles, and turnagra

Order: Passeriformes   Family: Oriolidae

The Old World orioles are colourful passerine birds. They are not related to the New World orioles.

Drongos
Black drongo

Order: Passeriformes   Family: Dicruridae

The drongos are mostly black or dark grey in colour, sometimes with metallic tints. They have long forked tails, and some Asian species have elaborate tail decorations. They have short legs and sit very upright when perched, like a shrike. They flycatch or take prey from the ground.


Fantails and silktails

Order: Passeriformes   Family: Rhipiduridae

The fantails are small insectivorous birds which are specialist aerial feeders.

Monarchs
Indian paradise flycatcher

Order: Passeriformes   Family: Monarchidae

The monarch flycatchers are small to medium-sized insectivorous passerines which hunt by flycatching.

Crows and jays

Order: Passeriformes   Family: Corvidae

The family Corvidae includes crows, ravens, jays, choughs, magpies, treepies, nutcrackers and ground jays. Corvids are above average in size among the Passeriformes, and some of the larger species show high levels of intelligence.

Tits and chickadees
Indian black-lored tit

Order: Passeriformes   Family: Paridae

The Paridae are mainly small stocky woodland species with short stout bills. Some have crests. They are adaptable birds, with a mixed diet including seeds and insects.

Larks
Ashy-crowned sparrow-lark

Order: Passeriformes   Family: Alaudidae

Larks are small terrestrial birds with often extravagant songs and display flights. Most larks are fairly dull in appearance. Their food is insects and seeds.

Bulbuls
Red-vented bulbul

Order: Passeriformes   Family: Pycnonotidae

Bulbuls are medium-sized songbirds. Some are colourful with yellow, red or orange vents, cheeks, throats or supercilia, but most are drab, with uniform olive-brown to black plumage. Some species have distinct crests.

Swallows and martins
Barn swallow

Order: Passeriformes   Family: Hirundinidae

The family Hirundinidae is adapted to aerial feeding. They have a slender streamlined body, long pointed wings and a short bill with a wide gape. The feet are adapted to perching rather than walking, and the front toes are partially joined at the base.

Leaf warblers and allies

Order: Passeriformes   Family: Phylloscopidae

Leaf warblers are a family of small insectivorous birds found mostly in Eurasia and ranging into Wallacea and Africa. The species are of various sizes, often green-plumaged above and yellow below, or more subdued with grayish-green to grayish-brown colors.

Reed warblers, Grauer’s warbler, and allies
Blyth's reed warbler

Order: Passeriformes   Family: Acrocephalidae

The members of this family are usually rather large for "warblers". Most are rather plain olivaceous brown above with much yellow to beige below. They are usually found in open woodland, reedbeds, or tall grass. The family occurs mostly in southern to western Eurasia and surroundings, but it also ranges far into the Pacific, with some species in Africa.

Grassbirds and allies

Order: Passeriformes   Family: Locustellidae

Locustellidae are a family of small insectivorous songbirds found mainly in Eurasia, Africa, and the Australian region. They are smallish birds with tails that are usually long and pointed, and tend to be drab brownish or buffy all over.

Cisticolas and allies
Plain prinia
Common tailorbird

Order: Passeriformes   Family: Cisticolidae

The Cisticolidae are warblers found mainly in warmer southern regions of the Old World. They are generally very small birds of drab brown or grey appearance found in open country such as grassland or scrub.

Sylviid babblers

Order: Passeriformes   Family: Sylviidae

The family Sylviidae is a group of small insectivorous passerine birds. They mainly occur as breeding species, as the common name implies, in Europe, Asia and, to a lesser extent, Africa. Most are of generally undistinguished appearance, but many have distinctive songs.

Parrotbills and allies
Yellow-eyed babbler

Order: Passeriformes   Family: Paradoxornithidae

The parrotbills are a group of peculiar birds native to East and Southeast Asia, though feral populations exist elsewhere. They are generally small, long-tailed birds which inhabit reedbeds and similar habitat. They feed mainly on seeds, e.g. of grasses, to which their bill, as the name implies, is well-adapted.

White-eyes
Indian white-eye

Order: Passeriformes   Family: Zosteropidae

The white-eyes are small and mostly undistinguished, their plumage above being generally some dull colour like greenish-olive, but some species have a white or bright yellow throat, breast or lower parts, and several have buff flanks. As their name suggests, many species have a white ring around each eye.

Babblers and scimitar babblers

Order: Passeriformes   Family: Timaliidae

The babblers, or timaliids, are somewhat diverse in size and colouration, but are characterised by soft fluffy plumage.

Ground babblers

Order: Passeriformes   Family: Pellorneidae

These small to medium-sized songbirds have soft fluffy plumage but are otherwise rather diverse. Members of the genus Illadopsis are found in forests, but some other genera are birds of scrublands.

Alcippe fulvettas
Brown-cheeked fulvetta or Quaker babbler

Order: Passeriformes   Family: Alcippeidae

The genus once included many other fulvettas and was previously placed in families Pellorneidae or Timaliidae.

Laughingthrushes and allies
Common babbler

Order: Passeriformes   Family: Leiothrichidae

The members of this family are diverse in size and colouration, though those of genus Turdoides tend to be brown or greyish. The family is found in Africa, India, and southeast Asia.

Nuthatches

Order: Passeriformes   Family: Sittidae

Nuthatches are small woodland birds. They have the unusual ability to climb down trees head first, unlike other birds which can only go upwards. Nuthatches have big heads, short tails and powerful bills and feet.

Treecreepers

Order: Passeriformes   Family: Certhiidae

Treecreepers are small woodland birds, brown above and white below. They have thin pointed down-curved bills, which they use to extricate insects from bark. They have stiff tail feathers, like woodpeckers, which they use to support themselves on vertical trees.

Starlings and rhabdornis
Brahminy starling

Order: Passeriformes   Family: Sturnidae

Starlings are small to medium-sized passerine birds. Their flight is strong and direct and they are very gregarious. Their preferred habitat is fairly open country. They eat insects and fruit. Plumage is typically dark with a metallic sheen.

Thrushes and allies

Order: Passeriformes   Family: Turdidae

The thrushes are a group of passerine birds that occur mainly in the Old World. They are plump, soft plumaged, small to medium-sized insectivores or sometimes omnivores, often feeding on the ground. Many have attractive songs.

Old World flycatchers
Indian robin
Asian brown flycatcher
Pied bushchat

Order: Passeriformes   Family: Muscicapidae

Old World flycatchers are a large group of small passerine birds native to the Old World. They are mainly small arboreal insectivores. The appearance of these birds is highly varied, but they mostly have weak songs and harsh calls.

Leafbirds
Golden-fronted leafbird

Order: Passeriformes   Family: Chloropseidae

The leafbirds are small, bulbul-like birds. The males are brightly plumaged, usually in greens and yellows.

Flowerpeckers

Order: Passeriformes   Family: Dicaeidae

The flowerpeckers are very small, stout, often brightly coloured birds, with short tails, short thick curved bills and tubular tongues.

Sunbirds
Crimson sunbird
Purple-rumped sunbird

Order: Passeriformes   Family: Nectariniidae

The sunbirds and spiderhunters are very small passerine birds which feed largely on nectar, although they will also take insects, especially when feeding young. Flight is fast and direct on their short wings. Most species can take nectar by hovering like a hummingbird, but usually perch to feed.

Old World sparrows and snowfinches

Order: Passeriformes   Family: Passeridae

Sparrows are small passerine birds. In general, sparrows tend to be small, plump, brown or grey birds with short tails and short powerful beaks. Sparrows are seed eaters, but they also consume small insects.

Weavers and widowbirds
Baya weaver

Order: Passeriformes   Family: Ploceidae

The weavers are small passerine birds related to the finches. They are seed-eating birds with rounded conical bills. The males of many species are brightly coloured, usually in red or yellow and black, some species show variation in colour only in the breeding season.

Waxbills, munias, and allies
Red avadavat

Order: Passeriformes   Family: Estrildidae

The estrildid finches are small passerine birds of the Old World tropics and Australasia. They are gregarious and often colonial seed eaters with short thick but pointed bills. They are all similar in structure and habits, but have wide variation in plumage colours and patterns.

Wagtails and pipits
White-browed wagtail
Paddyfield pipit

Order: Passeriformes   Family: Motacillidae

Motacillidae is a family of small passerine birds with medium to long tails. They include the wagtails, longclaws and pipits. They are slender, ground feeding insectivores of open country.

Finches and euphonias
Common rosefinch

Order: Passeriformes   Family: Fringillidae

Finches are seed-eating passerine birds, that are small to moderately large and have a strong beak, usually conical and in some species very large. All have twelve tail feathers and nine primaries. These birds have a bouncing flight with alternating bouts of flapping and gliding on closed wings, and most sing well.

Buntings

Order: Passeriformes   Family: Emberizidae

The emberizids are a large family of passerine birds. They are seed-eating birds with distinctively shaped bills. In Europe, most species are called buntings. In North America, most of the species in this family are known as sparrows, but these birds are not closely related to the Old World sparrows which are in the family Passeridae. Many emberizid species have distinctive head patterns.

See also

References
  1. "Telangana Declared Its Official Birds And Trees". SanctuariesIndia.com. 28 November 2014. Retrieved 10 January 2015.
  2. Kalinadhabhatla, A (2013). "Indian White-rumped Spinetail from Maredumilli, Andhra Pradesh". Indian Birds. 8 (6): 165.
  3. Jønsson, K.A., Bowie, R.C.K., Moyle, R.G., Irestedt, M., Christidis, L., Norman, J.A. & Fjeldså, J. (2010). "Phylogeny and biogeography of Oriolidae (Aves: Passeriformes)" (PDF). Ecography. 33: 232–241. doi:10.1111/j.1600-0587.2010.06167.x.CS1 maint: multiple names: authors list (link)
  4. Packert, Martin; Jochen Martens, Siegfried Eck, Alexander A Nazarenko, Olga P. Valchuk, Bernd Petri, Michael Veith (2005) The great tit (Parus major) – a misclassified ring species. Biological Journal of the Linnean Society 86(2):153-174
  5. Silke Fregin; Martin Haase; Urban Olsson; Per Alström (2009). "Multi-locus phylogeny of the family Acrocephalidae (Aves: Passeriformes) – The traditional taxonomy overthrown". Molecular Phylogenetics and Evolution. 52 (3): 866–878. doi:10.1016/j.ympev.2009.04.006. PMID 19393746.
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