Vanda

Vanda, abbreviated in the horticultural trade as V.,[2] is a genus in the orchid family, Orchidaceae. There are about 80 species, and the genus is commonly cultivated for the marketplace. This genus and its allies are considered to be among the most specifically adapted of all orchids within the Orchidaceae. The genus is highly prized in horticulture for its showy, fragrant, long-lasting, and intensely colorful flowers.[3] Vanda species are widespread across East Asia, Southeast Asia, and New Guinea, with a few species extending into Queensland and some of the islands of the western Pacific.[1][4]

For other uses, see Vanda (disambiguation).

Vanda
Vanda coerulea
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Monocots
Order: Asparagales
Family: Orchidaceae
Subfamily: Epidendroideae
Tribe: Vandeae
Subtribe: Aeridinae
Genus: Vanda
Gaud. ex Pfitzer
Type species
Vanda tessellata
Synonyms[1]
  • Ascocentrum Schltr.
  • Euanthe Schltr.
  • Finetia Schltr. 1918, illegitimate homonym, not Gagnep. 1917
  • Neofinetia Hu
  • Nipponorchis Masam. illegitimate
  • Eparmatostigma Garay
  • Trudelia Garay
  • × Trudelianda Garay
  • Christensonia Haager
  • Ascocentropsis Senghas & Schildh.
  • Gunnaria S.C.Chen ex Z.J.Liu & L.J.Chen

Biology

The name "Vanda" is derived from the Sanskrit (वन्दाका)[5] name for the species Vanda roxburghii (a synonym of Vanda tessellata).[6][7]

These mostly epiphytic, but sometimes lithophytic or terrestrial orchids, are distributed in India, Himalaya, Southeast Asia, Indonesia, the Philippines, New Guinea, southern China, and northern Australia.

The genus has a monopodial growth habit with highly variable leaves according to habitat. Some have flat, typically broad, ovoid leaves (strap-leaves), while others have cylindrical (terete), fleshy leaves and are adapted to dry periods. The stems of these orchids vary considerably in size; some are miniature plants and some have a length of several meters. The plants can become quite massive in habitat and in cultivation, and epiphytic species possess very large, rambling aerial root systems.

The few to many flattened flowers grow on a lateral inflorescence. Most show a yellow-brown color with brown markings, but they also appear in white, green, orange, red, and burgundy shades. The lip has a small spur. Vanda species usually bloom every few months and the flowers last for two to three weeks.

Many Vanda orchids (especially V. coerulea) are endangered, and have never been common because they are usually only infrequently encountered in habitat and grow only in disturbed forest areas with high light levels, and are severely threatened and vulnerable to habitat destruction.[3] The export of wild-collected specimens of the blue orchid (V. coerulea) and other wild Vanda species is prohibited worldwide, as all orchids are listed on Appendix II of the Convention on International Trade in Endangered Species.

Cultivation

This genus is one of the five most horticulturally important orchid genera, because it has some of the most magnificent flowers to be found in the orchid family. This has contributed much to the work of hybridists producing flowers for the cut flower market. V. coerulea is one of the few botanical orchids which can produce varieties with blue flowers (actually a very bluish purple), a property much appreciated for producing interspecific and intergeneric hybrids.

The color blue is rare among orchids, and only Thelymitra crinita, a terrestrial species from Australia, produces flowers that are truly "blue" among the orchids, the other being Aganisia cyanea, a lowland species from northern South America that is difficult to cultivate, but has metallic blue flowers. Both of these species, much like Vanda, also have a bluish-purple tint towards the inner petals of the flowers.

V. dearei is one of the chief sources of yellow color in Vanda hybrids. Vanda 'Miss Joaquim', a terrestrial Vanda variety and natural hybrid with terete (cylindrical) leaves, is the national flower of Singapore.

The plants do not possess pseudobulbs, but do possess leathery, drought-resistant leaves. Almost all of the species in this genus are very large epiphytes found in disturbed areas in habitat and prefer very high light levels, the plants having large root systems. Some of these species have a monopodial vine-like growth habit, and the plants can quickly become quite massive.

These plants prefer consistent conditions day-to-day in cultivation to avoid dropping their bottom leaves. The epiphytic species are best accommodated in large wooden baskets, bare rooted, which allows for the large aerial root systems. Disturbing or damaging the roots of large, mature vandaceous orchid plants, and in particular, Vanda and Aerides species, can result in the plants failing to flower and going into decline for a season or more. These plants do not tolerate disturbance or damage of their root systems in cultivation when they become mature. The terete-leaved terrestrial species are very easy to cultivate.

When grown bare-rooted, the epiphytic species require daily watering and weekly feeding and are very heavy feeders in cultivation. They can be grown out-of-doors in Hawaii and the like provided they are given some shade.

Systematics

In a recent molecular study of the genus Vanda ,[8] several Genera including the former Genus Ascocentrum, Neofinetea and Euanthe were brought into synonym with Vanda,[9][10]

Species

The following is a list of Vanda species recognised by the World Checklist of Selected Plant Families as at January 2019:[11]

Vanda ampullacea
Vanda Robert's Delight
Vanda tricolor
Vanda falcata
Vanda hindsii by Lewis Roberts
Vanda lamellata
Vanda garayi
  • Vanda aliceae
  • Vanda alpina (Himalaya to China - S. Yunnan)
  • Vanda × amoena
  • Vanda ampullacea
  • Vanda arbuthnotiana (India)
  • Vanda arcuata (Indonesia - Sulawesi)
  • Vanda aurantiaca
    • Vanda aurantiaca subsp. aurantiaca
    • Vanda aurantiaca subsp. philippinensis
  • Vanda aurea
  • Vanda barnesii (North Luzon, Philippines)
  • Vanda bensonii (Assam to Thailand)
  • Vanda bicolor (Bhutan)
  • Vanda bidupensis (Vietnam)
  • Vanda × boumaniae
  • Vanda brunnea (China - Yunnan to Indo-China)
  • Vanda celebica (Indonesia – Sulawesi)
  • Vanda × charlesworthii
  • Vanda chlorosantha (Bhutan)
  • Vanda christensoniana
  • Vanda coerulea : "Blue Orchid" (Assam to China - S. Yunnan)
  • Vanda coerulescens (Arunachal Pradesh to China - S. Yunnan)
  • Vanda concolor (S. China to Vietnam)
  • Vanda × confusa
  • Vanda cootesii
  • Vanda cristata (Himalaya to China - NW. Yunnan)
  • Vanda curvifolia
  • Vanda dearei (Borneo)
  • Vanda denisoniana (China - Yunnan to N. Indo-China)
  • Vanda devoogtii (Sulawesi)
  • Vanda dives
  • Vanda falcata (Japan, China, Korea)
  • Vanda flabellata (Rolfe ex Downie) Christenson, 1985
  • Vanda foetida (S. Sumatra)
  • Vanda frankieana
  • Vanda funingensis
  • Vanda furva (Java, Maluku)
  • Vanda fuscoviridis (S. China to Vietnam)
  • Vanda garayi
  • Vanda gibbsiae
  • Vanda gracilis
  • Vanda griffithii (E. Himalaya)
  • Vanda hastifera (Borneo)
  • Vanda helvola (W. Malaysia to Philippines)
  • Vanda hindsii (Papuasia to N. Queensland)
  • Vanda insignis (Lesser Sunda Is.)
  • Vanda insularum
  • Vanda jainii (Assam)
  • Vanda javierae (Philippines - Luzon)
  • Vanda jennae P.O'Byrne & J.J.Verm., 2005
  • Vanda lamellata (Taiwan, Philippines, Sabah)
    • Vanda lamellata var. boxallii
    • Vanda lamellata var. lamellata
    • Vanda lamellata var. remediosae
  • Vanda lilacina (China - Yunnan to Indo-China)
  • Vanda limbata (Java, Lesser Sunda Is., Philippines - Mindanao)
  • Vanda liouvillei (Assam to Indo-China)
  • Vanda lombokensis (Lesser Sunda Is.)
  • Vanda longitepala
  • Vanda luzonica (Philippines - Luzon)
  • Vanda malipoensis
  • Vanda mariae
  • Vanda merrillii (Philippines)
  • Vanda metusalae P.O'Byrne & J.J.Verm. (2008)
  • Vanda mindanoensis
  • Vanda miniata
  • Vanda motesiana
  • Vanda nana
  • Vanda perplexa
  • Vanda petersiana (Myanmar)
  • Vanda pumila (Nepal to Hainan and N. Sumatra)
  • Vanda punctata (Pen. Malaysia)
  • Vanda richardsiana
  • Vanda roeblingiana (Philippines - Luzon)
  • Vanda rubra
  • Vanda sanderiana
  • Vanda saxatilis
  • Vanda scandens (Borneo, Philippines - Mindanao)
  • Vanda semiteretifolia
  • Vanda stangeana (India - Arunachal Pradesh to Assam)
  • Vanda subconcolor (China - SW. Yunnan to Hainan)
  • Vanda sumatrana (Sumatra)
  • Vanda tessellata (Indian subcontinent to Indo-China)
  • Vanda testacea (Indian subcontinent to SC. China).
  • Vanda thwaitesii (S. India, Sri Lanka)
  • Vanda tricolor (Laos, Java, Bali)
    • Vanda tricolor var. suavis
    • Vanda tricolor var. tricolor
  • Vanda ustii (Philippines - Luzon)
  • Vanda vietnamica
  • Vanda vipanii (Myanmar)
  • Vanda wightii (S. India)
  • Vanda xichangensis

Natural hybrids
  • Vanda × boumaniae (V. insignis × V. limbata) (Lesser Sunda Is.)
  • Vanda × charlesworthii (V. bensonii × V. coerulea) (Myanmar)
  • Vanda × confusa (V. coerulescens × V. lilacina) (Myanmar)

Intergeneric hybrids
Vanda 'Pachara Delight'
Vanda Robert's Delight' × 'Crownfox Magic'
Vanda 'Sansai Blue'

The following is a list of hybrid genera (nothogenera) in which hybrids vandas with orchids of other genera are placed:

  • Aeridovanda (Aerides × Vanda)
  • Aeridovanisia (Aerides × Luisia × Vanda)
  • Alphonsoara (Arachnis × Ascocentrum × Vanda × Vandopsis)
  • Andrewara (Arachnis × Renanthera × Trichoglottis × Vanda)
  • Aranda (Arachnis × Vanda)
  • Ascocenda (Ascocentrum × Vanda)
  • Ascovandoritis (Ascocentrum × Doritis × Vanda)
  • Bokchoonara (Arachnis × Ascocentrum × Phalaenopsis × Vanda)
  • Bovornara (Arachnis × Ascocentrum × Rhynchostylis × Vanda)
  • Burkillara (Aerides × Arachnis × Vanda)
  • Charlieara (Rhynchostylis × Vanda × Vandopsis)
  • Christieara (Aerides × Ascocentrum × Vanda)
  • Darwinara (Ascocentrum × Neofinetia × Rhynchostylis × Vanda)
  • Debruyneara (Ascocentrum × Luisia × Vanda)
  • Devereuxara (Ascocentrum × Phalaenopsis × Vanda)
  • Eastonara (Ascocentrum × Gastrochilus × Vanda)
  • Fujiora (Ascocentrum × Trichoglottis × Vanda)
  • Goffara (Luisia × Rhynchostylis × Vanda)
  • Hawaiiara (Renanthera × Vanda × Vandopsis)
  • Hagerara (Doritis × Phalaenopsis × Vanda)
  • Himoriara (Ascocentrum × Phalaenopsis × Rhynchostylis × Vanda)
  • Holttumara (Arachnis × Renanthera × Vanda)
  • Isaoara (Aerides × Ascocentrum × Phalaenopsis × Vanda)
  • Joannara (Renanthera × Rhynchostylis × Vanda)
  • Kagawara (Ascocentrum × Renanthera × Vanda)
  • Knappara (Ascocentrum × Rhynchostylis × Vanda × Vandopsis)
  • Knudsonara (Ascocentrum × Neofinetia × Renanthera × Rhynchostylis × Vanda)
  • Leeara (Arachnis × Vanda × Vandopsis)
  • Luisanda (Luisia × Vanda)
  • Luivanetia (Luisia × Neofinetia × Vanda)
  • Lewisara (Aerides × Arachnis × Ascocentrum × Vanda)
  • Maccoyara (Aerides × Vanda × Vandopsis)
  • Macekara (Arachnis × Phalaenopsis × Renanthera × Vanda × Vandopsis)
  • Micholitzara (Aerides × Ascocentrum × Neofinetia × Vanda)
  • Moirara (Phalaenopsis × Renanthera × Vanda)
  • Mokara (Arachnis × Ascocentrum × Vanda)
  • Nakamotoara (Ascocentrum × Neofinetia × Vanda)
  • Nobleara (Aerides × Renanthera × Vanda)
  • Okaara (Ascocentrum × Renanthera × Rhynchostylis × Vanda)
  • Onoara (Ascocentrum × Renanthera × Vanda × Vandopsis)
  • Opsisanda (Vanda × Vandopsis)
  • Pageara (Ascocentrum × Luisia × Rhynchostylis × Vanda)
  • Pantapaara (Ascoglossum × Renanthera × Vanda)
  • Paulara (Ascocentrum × Doritis × Phalaenopsis × Renanthera × Vanda)
  • Pehara (Aerides × Arachnis × Vanda × Vandopsis)
  • Pereiraara (Aerides × Rhynchostylis × Vanda)
  • Phalaerianda (Aerides × Phalaenopsis × Vanda)
  • Raganara (Renanthera × Trichoglottis × Vanda)
  • Ramasamyara (Arachnis × Rhynchostylis × Vanda)
  • Renafinanda (Neofinetia × Renanthera × Vanda)
  • Renanda (Arachnis × Renanthera × Vanda)
  • Renantanda (Renanthera × Vanda)
  • Rhynchovanda (Rhynchostylis × Vanda)
  • Ridleyare (Arachnis × Trichoglottis × Vanda)
  • Robinaria (Aerides × Ascocentrum × Renanthera × Vanda)
  • Ronnyara (Aerides × Ascocentrum × Rhynchostylis × Vanda)
  • Sanjumeara (Aerides × Neofinetia × Rhynchostylis × Vanda)
  • Sarcovanda (Sarcochilus × Vanda)
  • Shigeuraara (Ascocentrum × Ascoglossum × Renanthera × Vanda)
  • Stamariaara (Ascocentrum × Phalaenopsis × Renanthera × Vanda)
  • Sutingara (Arachnis × Ascocentrum × Phalaenopsis × Vanda × Vandopsis)
  • Teohara (Arachnis × Renanthera × Vanda × Vandopsis)
  • Trevorara (Arachnis × Phalaenopsis × Vanda)
  • Trichovanda (Trichoglottis × Vanda)
  • Vascostylis (Ascocentrum × Rhynchostylis × Vanda)
  • Vandachnis (Arachnis × Vandopsis)
  • Vancampe (Acampe × Vanda)
  • Vandachostylis (Rhynchostylis × Vanda)
  • Vandaenopsis (Phalaenopsis × Vanda)
  • Vandaeranthes (Aeranthes × Vanda)
  • Vandewegheara (Ascocentrum × Doritis × Phalaenopsis × Vanda)
  • Vandofinetia (Neofinetia × Vanda)
  • Vandofinides (Aerides × Neofinetia × Vanda)
  • Vandoritis (Doritis × Vanda)
  • Vanglossum (Ascoglossum × Vanda)
  • Wilkinsara (Ascocentrum × Vanda × Vandopsis)
  • Yapara (Phalaenopsis × Rhynchostylis × Vanda)
  • Yusofara (Arachnis × Ascocentrum × Renanthera × Vanda)
  • Yonezawaara (Neofinetia × Rhynchostylis × Vanda)

References
  1. Kew World Checklist of Selected Plant Families
  2. "Alphabetical list of standard abbreviations of all generic names occurring in current use in orchid hybrid registration as at 31st December 2007" (PDF). Royal Horticultural Society.
  3. The Orchids, Natural History and Classification, Robert L. Dressler. ISBN 0-674-87526-5
  4. Flora of China v 25 p 471, 万代兰属 wan dai lan shu, Vanda Jones ex R. Brown, Bot. Reg. 6: ad t. 506. 1820.
  5. vandAkA Sanskrit English Dictionary, University of Koeln, Germany
  6. Vanda Taxonomy Research & Information Network, Australia (2009)
  7. Garay, L. (1972), On the systematics of the monopodial orchids, Bot. Mus. Leafl. Harvard University, 23(4): 149-212
  8. Lim, S. (1999). "RAPD Analysis of Some Species in the GenusVanda(Orchidaceae)". Annals of Botany. 83 (2): 193–196. doi:10.1006/anbo.1998.0801.
  9. http://www.ipni.org/ipni/idPlantNameSearch.do?id=661063-1&back_page=%2Fipni%2FeditSimplePlantNameSearch.do%3Ffind_wholeName%3DVanda%2Bsanderiana%26output_format%3Dnormal
  10. http://apps.kew.org/wcsp/namedetail.do?name_id=211529M
  11. World Checklist of Selected Plant Families : Vanda

Further reading
  • Grove, D. L. 1995. Vandas and Ascocendas. Timber Press, Portland, Oregon. 241 pp.
  • Motes, Martin R., and Alan L. Hoffman. 1997 Vandas, Their botany, history and culture. ISBN 0-88192-376-1
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