Coleoptera paleobiota of Burmese amber

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018;[1] its supplement Ross 2019b covers most of 2019.[2]

This article covers fossils classified as belonging to order Coleoptera, and its sub-orders Adephaga, Archostemata, Myxophaga, and Polyphaga.

Adephaga

Taxon	Authority	Year described	Notes
Ambarticus myanmaricus[3]	Yang, Chen & Jia	2019	A diving beetle.
Burmapseudomorphus planus[4]	Beutel et al.	2020	A pseudomorphine ground beetle.
Chimerogyrus gigagalea[5]	Gustafson, Michat & Balke	2020	A gyrinid whirligig beetle
Cretodineutus rotundus[6]	Liang et al.	2020	A gyrinine whirligig beetle
Cretogyrus beuteli[7]	Zhao et al.	2019	A whirligig beetle.
Kryzhanovskiana olegi[8]	Kataev et al.	2019	A Metriini ant nest beetle.

Archostemata

Taxon	Authority	Year described	Notes
Barbaticupes combertiae[9]	Jarzembowski, Wang & Zheng	2017	A reticulated beetle.
Bukhkalius lindae[10]	(Jarzembowski, Wang & Zheng)	(2017)	An ommatine beetle. The species was first described as "Tetraphalerus" lindae.
Burmocoleus prisnyi[10]	Kirejtshuk	2020	An ommatine beetle. "Brochocoleus" zhiyuani (2017).
Burmocoleus zhiyuani[10]	(Liu et al.)	2017	An ommatine beetle. first described as "Brochocoleus" zhiyuani.[11]
Cionocups manukyani[10]	Kirejtshuk	2020	An ommatine beetle.
Clessidromma palmeri[12]	Jarzembowski, Wang & Zheng	2017	An Ommatine beetle.
Clessidromma tianae[13]	(Jarzembowski, Wang & Zheng)	2019	An ommatine beetle. Originally described as "Lepidomma" tianae.[10]
Clessidromma zengi[10]	Kirejtshuk	2020	An ommatine beetle.
Echinocups denticollis[14][10]	(Jiang et al.)	2020	An ommatine beetle. Originally described as "Notocupes" denticollis.
Echinocups neli[10]	(Tihelka, Huang & Cai)	2010	An ommatine. Originally described as "Notocupes" neli.
Echinocups ohmkuhnlei[15][10]	(Jarzembowski, Wang & Zheng)	2020	An ommatine beetle. Originally described as "Notocupes" ohmkuhnlei.
Jarzembowskiops caseyi[10]	(Jarzembowski, Wang & Zheng)	2020	An ommatine beetle. First described as "Brochocoleus" caseyi.
Lepidomma jarzembowskii[16]	Li et al.	2020	An ommatine beetle.
Lepidomma longisquama[16]	Li et al.	2020	An ommatine beetle.
Mallecupes cleevelyi[17]	Jarzembowski, Wang & Zheng	2017	A reticulated beetle.
Mallecupes prokini[10]	Kirejtshuk	2020	A reticulated beetle,
Mallecupes qingqingae[18]	Jarzembowski, Wang & Zheng	2016	A reticulated beetle.
Miniomma chenkuni[19]	Li, Yamamoto & Cai in Li et al.	2020	An ommatid beetle.
Notocupes neli[20]	Tihelka, Huang & Cai	2019	A reticulated beetle.
Omma davidbatteni[21]	Jarzembowski, Wang & Zheng	2020	An ommatid beetle.
Omma janetae[10]	Kirejtshuk	2020	An ommatid beetle.
Omma lii[22]	Jarzembowski, Wang & Zheng	2017	An ommatid beetle.
Paraodontomma burmitica[23]	Yamamoto	2017	A Brochocoleini Ommatid beetle.
Paraodontomma szwedoi[24]	Jarzembowski, Wang & Zheng	2018	A Brochocoleini Ommatid beetle.
Priacma megapuncta[25]	Li et al.	2019	A reticulated beetle.
Protomalthus burmaticus[26]	Tihelka, Huang & Cai	2019	A telephone-pole beetle.
Polyakius alberti[10]	Kirejtshuk	2020	An ommatine beetle.
Polyakius pubescens[10]	Kirejtshuk	2020	An ommatine beetle.
Stegocoleus arkonus[27]	Tihelka, Huang & Cai	2020	An ommatid beetle.
Stegocoleus caii[28]	Jarzembowski & Wang	2016	An ommatid beetle.
Stegocoleus lawrencei[27]	Tihelka, Huang & Cai	2020	An ommatid beetle.

Myxophaga

Taxon	Authority	Year described	Notes
Burmasporum rossi[29]	Kirejtshuk	2009	A sphaeriusid beetle.
Lepiceratus ankylosaurus[30]	Jałoszyński et al.	2020	A stem group member of the family Lepiceridae.
Lepicerus mumia[31]	Jałoszyński & Yamamoto	2017	A Lepicerid beetle.
Lepichelus pretiosus[32]	(Kirejtshuk & Poinar)	2013	A Lepicerid beetle, Originally described as "Lepiceroides" pretiosus.[33]

Polyphaga

Taxon	Authority	Year described	Notes
Acalyptomerus thayerae[34]	Cai & Lawrence in Cai et al.	2019	A fringe-winged beetle.
Acalyptopygus astriatus[35]	Clarke & Oberprieler in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil.
Acalyptopygus brevicornis[35]	Clarke & Oberprieler in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil.
Acalyptopygus elongatus[35]	Clarke & Oberprieler in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil.
Acalyptopygus lingziae[35]	Clarke & Oberprieler in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil.
"Acmaeodera" burmitina[36]	Cockerell	1917	An elaterid click beetle.
Adamas hukawngensis[37]	Chen & Zhang	2020	A ship-timber beetle.
Aepyceratus hyperochus[38]	Poinar, Brown & Legalov	2017	An Aepyceratinae mesophyletid weevil.[35]
Allostrophus cretaceus[39]	Hsiao et al.	2018	A polypore fungus beetle.
Allostrophus yangi[40]	Hsiao	2019	A polypore fungus beetle.
Amberathyreus beuteli[41]	Bai & Zhang	2017	An athyreine dor beetle
Amberocula costata[42]	Batelka, Engel, & Prokop	2018	A wedge-shaped beetle.
Amberocula fallax[42]	Batelka, Engel, & Prokop	2018	A wedge-shaped beetle.
Amberocula muelleri[42]	Batelka, Engel, & Prokop	2018	A wedge-shaped beetle.
Amberophytum birmanicum[43]	Yu, Ślipiński & Pang in Yu et al.	2019	A rare click beetle.
Amplectister tenax[44]	Caterino & Maddison	2018	A histerid clown beetle.
Anchineus dolichobothris[45]	Poinar & Brown	2009	A Mesophyletinae mesophyletid weevil.
Anebomorpha cercorhampha[46]	Poinar & Brown	2020	A clerid checkered beetle.
Angimordella burmitina[47]	Bao et al.	2019	A tumbling flower beetle.
Aphebodactyla rhetine[48]	Chatzimanolis, Cashion, Engel & Falin	2012	A toe-winged beetle.
Aphelonyssus latus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Apophisandra ammytae[49]	Molino-Olmedo	2017	A parandrine longhorn beetle.
Apotomoura fortiscrura[50]	Bao et al.	2018	An apotomourid or mordellid[51] beetle.
Archaeodraena cretacea[52]	Jäch & Yamamoto	2017	A minute moss beetle.
Archaeomalthodes rosetta[53]	Hsiao et al.	2017	A malthinine soldier beetle
Archaeozenodosus bellus[54]	Yu & Kolibač	2017	A thaneroclerid checkered beetle.
Archemastax divida[55]	Yin et al.	2019	An ant-like stone beetle.
Attagenus burmiticus[56]	Cai, Háva & Huang	2017	An attagenine carpet beetle.
Attagenus lundi[57]	Háva & Damgaard	2017	An attagenine carpet beetle.
Attagenus secundus[58]	Deng et al.	2017	An attagenine carpet beetle.
Basinosa pengweii[59]	Tihelka, Huang & Cai	2020	A wounded-tree beetle.
Bifurderum donoghuei[60]	Cheng et al.	2020	A cylindrical bark beetle.
Bifurderum minutum[60]	Cheng et al.	2020	A cylindrical bark beetle.
Boreotethys arctopteryx[61]	Parker	2016	A Pselaphinae rove beetle.
Boreotethys grimaldii[61]	Parker	2016	A Pselaphinae rove beetle.
Bowangius cyclops[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Bowangius glabratus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Bowangius tanaops[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Bowangius zhenuai[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Brachycerophytum cretaceum[43]	Yu, Ślipiński & Pang	2019	A rare click beetle.
Burmacateres longicoxa[62]	Kolibáč & Peris	2020	A lophocaterid beetle.
Burmahelota pengweii[63]	Liu et al.	2019	A helotid beetle.
Burmalestes albertalleni[64]	Tomaszewska & Ślipiński in Tomaszewska et al.	2018	An endomychid handsome fungus beetle.
Burmitoma nalae[42]	Batelka, Engel & Prokop	2018	A wedge-shaped beetle.
Burmocorynus jarzembowskii[65]	Legalov	2018	A Mesophyletinae mesophyletid weevil.[35]
Burmocorynus longus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Burmolycus compactus[66]	Bocak, Li & Ellenberger	2019	A dexorine net-winged beetle.
Burmomacer kirejtshuki[65]	Legalov	2018	A Nemonychidae weevil.
Burmomiles blixenae[67]	Fanti & Damgaard	2019	A soldier beetle
Burmomiles willerslevorum[68]	Fanti, Damgaard & Ellenberger	2018	A soldier beetle.
Burmonyx zigrasi[35]	David & Engel	2014	A nemonychid weevil
Burmophyletis beloides[35]	(Clarke & Oberprieler) in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil. Originally described as "Platychirus" beloides.[69]
Burmops neli[70]	Kirejtshuk, Willig & Chetverikov	2019	A protosphindine sphindid beetle.
Burmorhinus georgei[71]	Legalov	2018	A Mesophyletinae mesophyletid weevil.[35]
Burmorhinus setosus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Calyptocis brevirostris[35]	Clarke & Oberprieler in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil.
Captopus depressiceps[72]	Li & Cai in Li, Huang & Cai	2021	A throscid beetle.
Cascomastigus minor[73]	Yin, Cai & Huang	2018	A Scydmaeninae rove beetle.
Cedasyrus dimorphus[74]	Yin & Cai in Yin et al.	2020	A dasycerine rove beetle.
Cenomana clavata[75]	Otto	2019	A jenibuntorine macraulacine false click beetle.
Cenomaniola carinata[76]	Jałoszyński & Yamamoto	2017	A glandulariine ant-like stone beetle.
Cenomaniola macrophthalma[76]	Jałoszyński & Yamamoto	2017	A glandulariine ant-like stone beetle.
Ceracyclus jirouxi[77]	Boucher, Bai & Montreui	2017	A ceracycline aulacocycline bess beetle.
Ceracyclus lotus[77]	Boucher, Bai & Montreui	2017	A ceracycline aulacocycline bess beetle.
Cetionyx batiatus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Cetionyx terebrans[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Cetionyx ursinus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Cerophytum albertalleni[43]	Yu, Ślipiński & Pang	2019	A rare click beetle.
Clidicostigus arachnipes[78]	Jałoszyński, Brunke & Bai	2017	A mastigine ant-like stone beetle.
Clidicus archaicus[79]	(Cai & Huang)	2016	A scydmaenine rove beetle. First published as "Cretoleptochromus" archaicus.[80]
Coenomana brevicornis[81]	Muona	2020	A melasine false click beetle.
Colon burmiticum[82]	Yamamoto in Yamamoto & Takahashi	2018	A Coloninae round fungus beetle.
Colonellus burmiticus[83]	Cai & Huang	2017	A Coloninae round fungus beetle.
Compsopsarus reneae[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Cretaceoripidius burmiticus[42]	(Cockerell)	1917	A ripidiini ripidiine wedge shaped beetle. Originally described as "Myodites" burmiticus, moved from "Ripiphorus" burmiticus.[84]
Cretagyrtodes glabratus[85]	Cai & Huang	2017	An agyrtodine Camiarinae round fungus beetle is C. .
Cretakarenni birmanicus[86]	Peris & Delclòs	2015	A monotomid beetle.
Cretaparamecus tarsalis[64]	Tomaszewska, Ślipiński, Bai & Zhang in Tomaszewska et al.	2018	A handsome fungus beetle.
Cretaretes minimus[87]	Peris & Jelínek	2020	A kateretid short-winged flower beetle
Cretasernus spinosus[88]	Peris et al.	2019	A ptinid beetle.
Cretoattagenus coziki[89]	Háva	2020	An attagenine carpet beetle.
Cretoboganium gei[90]	Cai & Huang	2019	A boganiine beetle.
Cretobrachygluta laurasiensis[91]	Yin et al.	2019	A brachyglutine pselaphine rove beetle.
Cretobythus excavatus[92]	Yin, Parker & Cai	2017	A stem-bythinine pselaphine rove beetle.
Cretochirus newtoni[93]	Yamamoto	2019	A leptochirine osoriine rove beetle.
Cretocrenis burmanicus[94]	Fikáček et al.	2017	A water scavenger beetle.
Cretodeinopsis aenigmatica[95]	Cai & Huang	2015	An aleocharine rove beetle.
Cretodermestes palpalis[58]	Deng et al.	2017	An attagenine carpet beetle.
Cretohlezkus alleni[96]	Jałoszyński	2019	A plate-thigh beetle.
Cretohypsilara parva[97]	Cai, Maier & Huang	2018	A riffle beetle.
Cretolenax carinatus[98]	Liu, Tihelka, McElrath & Yamamoto	2020	A monotomid minute clubbed beetle
Cretolenax diabolus[98]	Tihelka, Liu, McElrath & Yamamoto	2020	A monotomid minute clubbed beetle
Cretoleptochromus burmiticus[99]	Yin et al.	2017	An ant-like stone beetle.
Cretolestes niger[64]	Tomaszewska, Ślipiński & Ren in Tomaszewska et al.	2018	A handsome fungus beetle.
Cretolgus minimus[100]	Legalov & Háva	2020	A polycaonine auger beetle.
Cretoliota cornutus[101]	Liu et al.	2019	A silvan flat bark beetle.
Cretolycus praecursor[102]	Tihelka, Huang & Cai	2019	A net-winged beetle.
Cretomysteria burmanica[103]	Denget al.	2017	A colydiine ironclad beetle.
Cretonthophilus tuberculatus[104]	Caterino, Wolf-Schwenninger & Bechly	215	A clown beetle.
Cretoparacucujus cycadophilus[105]	Cai & Escalona in Cai et al.	2018	A Paracucujinae boganiid beetle.
Cretophengodes azari[106]	Li, Kundrata, Tihelka & Cai in Cai et al.	2021	A member of Elateroidea belonging to the family Cretophengodidae.
Cretopityobius pankowskiorum[75]	Otto	2019	A pityobiine click beetle.
Cretoprocirrus trichotos[107]	Jenkins Shaw & Żyła in Jenkins Shaw et al.	2020	A Pinophilini rove beetle.
Cretopsectra pulchra[108]	Zhao et al.	2020	A brachypsectrid texas beetle.
Cretopseudopsis maweii[109]	Liu et al.	2020	A pseudopsine rove beetle.
Cretoptomaphagus microsoma[110]	Bao & Antunes-Carvalho	2020	A cholevine round fungus beetle.
Cretoquadratus engeli[111]	Chen	2019	A ship-timber beetle.
Cretosynstrophus archaicus[112]	Cai, Hsiao & Huang	2016	A Eustrophinae polypore fungus beetle.
Cretotrichopsenius burmiticus[113]	Cai et al.	2017	A Trichopseniine Aleocharine rove beetle.
Cretozenodosus fossilis[114]	Cai & Huang	2018	A Zenodosinae thaneroclerid checkered beetle.
Cryphalites rugosissimus[115]	Cockerell	1917	A colydiine iron-clad beetle
Cupressicharis elongatus[81]	Muona	2020	A palaeoxenine false click beetle.
Cylus carinifer[81]	Muona	2020	A eucnemine false click beetle.
Cyrtocis gibbus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Debbia gracilirostris[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Dermestes larvalis[116]	Cockerell	1917	A dermestine Dermestinae Dermestid beetle
Derolathrus abyssus[117]	Yamamoto, Takahashi & Parker	2017	A Jacobsoniid beetle.
Diminudon kachinensis[118]	Żyła, Yamamoto & Shaw	2019	A paederine rove beetle
Diminudon schomannae[118]	Żyła, Yamamoto & Shaw	2019	A paederine rove beetle
Ditysparedrus gigas[119]	Vitali & Legalov	2020	A false blister beetle.
Ditysparedrus obscurus[119]	Vitali & Legalov	2020	A false blister beetle.
Ditysparedrus serixioides[119]	Vitali & Legalov	2020	A false blister beetle.
Ditysparedrus uniformis[119]	Vitali & Legalov	2020	A false blister beetle.
Echogomphus viridescens[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Ektatotricha paradoxa[120]	Chatzimanolis, Engel & Newton	2010	An ant-like stone beetle
"Elater" burmitinus[36]	Cockerell	1917	An elaterid click beetle.
Electraesalopsis beuteli[121]	Bai, Zhang & Qiu	2017	A stag beetle.
Electroatopos castaneus[120]	Chatzimanolis, Engel & Newton	2010	An ant-like stone beetle
Electrocis dentitibialis[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Electrorubesopsis beuteli[122]	Bai & Wang in Li et al.	2018	An Electrorubesopsinae scarab beetle.
Electrothroscus yanpingae[72]	Li & Cai in Li, Huang & Cai	2021	A throscid beetle.
Electroxyra cretacea[123]	(Wu, Li & Ding)	2018	A sooty mould beetle. Originally described as "Cyclaxyra" cretacea.[124]
Electrumeretes birmanicus[125]	Peris & Jelínek	2019	A short-winged flower beetle.
Elektrokleinia picta[126]	Ellenberger & Fanti	2019	A soldier beetle.
Elektrokleinia steffenseni[127]	Fanti & Damgaard	2020	A soldier beetle.
Eoceniretes antiquus[87]	Peris & Jelínek	2020	A kateretid short-winged flower beetle
Epiphanis burmensis[81]	Muona	2020	A species of Epiphanis.
Erichia cretacea[128]	Yu et al.	2018	A minute marsh-loving beetle.
Eucinetus parvus[129]	Du et al.	2020	A plate-thigh beetle.
Euryepomus lophomerus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
‘Eurygenius’ wickhami[36]	Cockerell	1917	A tenebrionoid beetle of uncertain placement.[130]
Euryptychus acutangulus[81]	Muona	2020	A macraulacine false click beetle.
Euryptychus burmensis[81]	Muona	2020	A macraulacine false click beetle.
Euryptychus elegantulus[81]	Muona	2020	A macraulacine false click beetle.
Euryptychus mysticus[81]	Muona	2020	A macraulacine false click beetle.
Excavotarsus lini[131]	Li et al.	2020	A heterocerid variegated mud-loving beetle.
Excavotarsus minor[131]	Li et al.	2020	A heterocerid variegated mud-loving beetle.
Falsothambus burmensis[81]	Muona	2020	A eucnemine false click beetle.
Falsothambus gracilicornis[81]	Muona	2020	A eucnemine false click beetle.
Festenus gracilis[132]	Żyła et al.	2017	A Stenine rove beetle.
Festenus robustus[132]	Żyła et al.	2017	A Stenine rove beetle.
Fiegelia antennata[81]	Muona	2020	A false click beetle.
Fiegelia tarsalis[81]	Muona	2020	A false click beetle.
Flabellotoma heidiae[133]	Batelka, Prokop & Engel	2016	A Pelecotominae wedge shaped beetle.
Furcalabratum burmanicum[134]	Poinar & Brown	2018	A short-winged flower beetle.
Glaresis burmitica[135]	Cai & Huang	2018	An enigmatic scarab beetle.
Gnomus brevis[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Gnomus spinipes[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Gollandia planata[136]	Makranczy, Yamamoto & Engel	2018	A Coprophilini Oxytelinae rove beetle.
Guillermorhinus longitarsis[35]	Clarke & Oberprieler in Clarke et al	2018	A nemonychid weevil
Habropezus kimpulleni[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Habropezus ncoxatirostris[35]	Clarke & Oberprieler in Clarke et al.		A Mesophyletinae mesophyletid weevil.
Habropezus plaisiommus[137]	Poinar, Brown & Legalov	2016	A Mesophyletinae mesophyletid weevil.[35]
Habropezus tenuicornis[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Hapsomela burmitis[138]	Poinar & Brown	2004	An ant-like stone beetle
Hapsomela minor[139]	Yin	2020	An ant-like stone beetle.
Hapsomela tibialis[139]	Yin	2020	An ant-like stone beetle.
Hongipsectra electrella[140]	Tihelka, Huang & Cai	2019	A Texas beetle.
Hukawngichthyurus kyawkhaingwini[141]	Fanti & Ellenberger	2018	A soldier beetle.
Hukawngius crassipes[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Hybosorus ocampoi[142]	Bai & Zhang in Bai et al.	2016	A scavenger scarab beetle.
Jenibuntor pusillus[81]	Muona	2020	A macraulacine false click beetle.
Kachinus antennatus[120]	Chatzimanolis, Engel & Newton	2010	An ant-like stone beetle
Kekveus jason[143]	Yamamoto, Grebennikov & Takahashi	2018	A featherwing beetle.
Kresnikus beynoni[144]	Tihelka, Huang & Cai	2019	A hide beetle.
Kuafu borealis[145]	Yin et al.	2018	A stem group Scydmaenini ant-like stone beetle.
Lemodicarmenia olmedoae[146]	Molino-Olmedo	2017	A Lemodine ant-like flower beetle.
Lenax karenae[98]	Liu, Tihelka, McElrath & Yamamoto	2020	A monotomid minute clubbed beetle
Leptopezus barbatus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae Mesophyletidae weevil.
Leptopezus rastellipes[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae Mesophyletidae weevil.
Loeblitoides separatus[147]	Jałoszyński	2020	A Glandulariini ant-like stone beetle.
Longicrusa jaracimrmani[148]	Tihelka, Huang & Cai	2019	A false darkling beetle.
Louwiocis megalops[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Megalopinus extinctus[149]	Yamamoto & Solodovnikov	2016	A Megalopsidiinae rove beetle.
Megatoma atypica[58]	Deng et al.	2017	A Megatomine carpet beetle.
Mekorhamphus beatae[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Mekorhamphus gracilipes[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Mekorhamphus gyralommus[137]	Poinar, Brown & Legalov	2016	A Mesophyletinae mesophyletid weevil.[35]
Mekorhamphus poinari[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Mekorhamphus tenuicornis[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Mesallotrochus longiantennatus[150]	Cai & Huang	2015	An osoriine rove beetle.
Mesernobius anawrahtai[151]	Engel	2010	A Mesernobiine marsh beetle
Mesolophocateres pengweii[152]	Yu, Leschen & Ślipiński in Yu et al.	2020	A lophocaterid beetle.
Mesopassandra keyao[153]	Jin et al.	2019	A mesopassandrine parasitic flat beetle.
Mesophyletis calhouni[154]	Poinar	2006	A Mesophyletinae mesophyletid weevil.
Mesosmicrips cretacea[155]	(Cai & Huang)	2016	A palmetto beetle. Originally described as "Smicrips" cretacea.[156]
Mesosymbion compactus[157]	Yamamoto, Maruyama & Parker	2016	A Mesoporini aleocharine rove beetle.
Metahelotella monochromata[63]	Liu et al.	2019	A helotid beetle.
Microborus inertus[158]	Cognato & Grimaldi	2009	A hexacoline scolytine weevil.
Microentomus epibatus[159]	Poinar in Poinar & Brown	2014	A blister beetle.
Micropeplus pengweii[160]	Jiang, Peng & Wang	2020	A micropepline rove beetle.
Molinernobius fuentesi[161]	Molino-Olmedo	2017	An ernobiine ptinid beetle.
Multispinus multispinosus[50]	Bao et al.	2018	An apotomourid or mordellid beetle.
Multispinus parvus[51]	Bao	2020	An apotomourid or mordellid beetle.
Muonabuntor grandinotalis[162]	Li et al.	2020	A false click beetle.
Myall burmensis[81]	Muona	2020	A eucnemine false click beetle.
Myamalycocerus vitalii[163]	Fanti & Ellenberger	2017	A soldier beetle.
Myanmarops gatiosus[164]	Legalov, Kirejtshuk & Anokhin	2020	A bruchine seed beetle
Myanmarus caviventris[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Myanmarus dentifer[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Myanmarus diversiunguis[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Myanmarus robustus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Mysteriomorphus pelevini[165]	Alekseev & Ellenberger	2019	An elateroid[166] mysteriomorphid beetle.
Necromeropsis minutus[43]	Yu, Ślipiński & Pang	2019	A rare click beetle.
Nosodendron cretaceum[167]	Deng et al.	2019	A wounded-tree beetle.
Nuegua elongata[168]	Yin, Cai & Newton	2018	A glandulariine ant-like stone beetle.
Nugatorhinus albomaculatus[35]	Clarke & Oberprieler in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil.
Nugatorhinus chenyangi[35]	Clarke & Oberprieler in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil.
Ocriocis binodosus[35]	Clarke & Oberprieler in Clarke et al.		A Mesophyletinae mesophyletid weevil.
Octavius electrospinosus[169]	Clarke & Chatzimanolis	2009	A Euaesthetine Euaesthetinae rove beetle
Onthophilus yingae[170]	Jiang et al.	2020	A clown beetle.
Oodes kachinensis[171]	Liu	2014	A ground beetle.
Opeatorhynchus comans[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Ornatomalthinus elvirae[172]	Poinar & Fanti	2016	A soldier beetle.
Oxyporus cretaceous[173]	Yamamoto	2017	An oxyporine rove beetle.
Pactopus burmensis[174]	Muona	2019	A throscid false click beetle.
Palaeocryptorhynchus burmanus[175]	Poinar	2009	A palaeocryptorhynchine weevil.
Palaeomycetes foveolatus[64]	Tomaszewska, Ślipiński & Ren in Tomaszewska et al.	2018	A handsome fungus beetle.
Paleoendeitoma antennata[103]	Denget al.	2017	A synchitine colydiine ironclad beetle.
Paleoendeitoma buryi[176]	Háva	2019	A synchitine colydiine ironclad beetle.
Paleoendeitoma minuta[103]	Denget al.	2017	A synchitine colydiine ironclad beetle.
Paleoendeitoma tuberculata[177]	Bullis	2020	A colydiinae cylindrical bark beetle.
Paleoeucnemis minutus[81]	Muona	2020	A eucnemine false click beetle.
Pangusyndicus excavatus[178]	Yin, Zhou & Cai in Yin et al.	2018	An ant-like stone beetle.
Pangusyndicus longirostris[147]	Jałoszyński	2020	A glandulariine ant-like stone beetle.
Pantostictus burmanicus[179]	Poinar & Brown	2009	A clown beetle
Parayixianteres parvus[152]	Yu, Leschen & Ślipiński in Yu et al.	2020	A lophocaterid beetle.
Passalopalpus cheni[180]	Boucher et al.	2016	A passalopalpid beetle.
Periosocerus crenulatus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Periosocerus deplanatus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Periosomerus tanyorhynchus[181]	Poinar, Brown & Legalov	2019	A New York weevil.
Petalotarsus curculionoides[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Petalotarsus cylindricus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Petalotarsus oxycorynoides[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Plesiotoma alissae[42]	Batelka, Engel, & Prokop	2018	A wedge-shaped beetle.
Pleuroceratos burmiticus[182]	Poinar & Kirejtshuk	2008	A phloeostichid beetle.
Pleuroceratos jiewenae[183]	Tihelka, Huang & Cai	2020	A phloeostichid beetle.
Poinarelektronmiles ellenbergeri[127]	Fanti & Damgaard	2020	A soldier beetle.
Poinarinius burmaensis[184]	Legalov	2018	A Dinoderinae auger beetle.
Polliniretes penalveri[125]	Peris & Jelínek	2019	A short-winged flower beetle.
Praezolodinus pilosus[110]	Bao & Antunes-Carvalho	2020	A zolodinine darkling beetle.
Prajna tianmiaoae[185]	Lü, Cai & Huang	2017	A thinobiine oxyteline rove beetle.
Praphennium carinatum[186]	Jałoszyński	2018	A Cephenniini Scydmaeninae rove beetle.
Primaevomordellida burmitina[187]	Bao et al.	2019	A tumbling flower beetle.
Priochirus thayerae[93]	Yamamoto	2019	An osoriine rove beetle.
Priscaplectus carinatus[188]	Yin, Chandler & Cai	2019	A pselaphine rove beetle
Priscaplectus grandiceps[188]	Yin, Chandler & Cai	2019	A pselaphine rove beetle.
Procileoporus burmiticus[189]	Yamamoto	2016	A Tachyporinae rove beetle.
Promyrmister kistneri[190]	Zhou, Ślipiński & Parker	2019	A haeteriine clown beetle.
Propiestus archaicus[191]	Yamamoto, Caron & Bortoluzzi	2018	A Piestinae rove beetle.
Prosolierius crassicornis[192]	Thayer, Newton & Chatzimanolis	2012	A solieriine rove beetle
Prosolierius mixticornis[192]	Thayer, Newton & Chatzimanolis	2012	A solieriine rove beetle
Prosolierius tenuicornis[192]	Thayer, Newton & Chatzimanolis	2012	A solieriine rove beetle
Protodasycerus aenigmaticus[193]	Yamamoto	2016	A dasycerine rove beetle.
Protodasycerus corpulentus[74]	Yin & Cai in Yin et al.	2020	A dasycerine rove beetle.
Protodasycerus gigas[74]	Yin & Cai in Yin et al.	2020	A dasycerine rove beetle.
Protodasycerus tuberculatus[74]	Yamamoto, Newton & Yin in Yin et al.	2020	A dasycerine rove beetle.
Protoliota antennatus[101]	Liu et al.	2019	A silvan flat bark beetle.
Protoluciola albertalleni[194]	Kazantsev	2015	A lucioline firefly.
Protomicrorhagus antennatus[81]	Muona	2020	A melasine false click beetle.
Protomicrorhagus brevis[81]	Muona	2020	A melasine false click beetle.
Protonicagus tani[195]	Cai et al.	2017	A nicagine aesaline stag beetle.
Protopeplus cretaceus[196]	Cai & Huang	2014	A micropepline rove beetle
Protopselaphus newtoni[197]	Liu et al.	2020	A protopselaphine rove beetle.
Protopselaphus thayerae[198]	Liu et al.	2020	A protopselaphine rove beetle.
Protoripidius burmiticus[199]	Cai, Yin & Huang	2018	A Ripidiinae wedge-shaped beetle.
Prototrichalus meiyingae[200]	Molino-Olmedo et al.	2020	A lycid net-winged beetle.
Prototrichalus milleri[200]	Molino-Olmedo et al.	2020	A lycid net-winged beetle.
Prototrichalus sepronai[200]	Molino-Olmedo et al.	2020	A lycid net-winged beetle.
Protovitellius deceptus[81]	Muona	2020	A eucnemine false click beetle.
Protrichonyx rafifrons[61]	Parker	2016	A Pselaphinae rove beetle.
Pseudomyall elongatulus[81]	Muona	2020	A eucnemine false click beetle.
Pseudopactopus robustus[72]	Li & Cai in Li, Huang & Cai	2021	A throscid beetle.
Qitianniu zhihaoi[201]	Lin & Bai	2017	A longhorn beetle.
Raractocetus extinctus[202]	Yamamoto	2019	An Atractocerinae ship-timber beetle.
Rhadinomycter perplexus[35]	Clarke & Oberprieler in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil.
Rhynchitomimus chalybeus[35]	Clarke & Oberprieler in Clarke et al.	2018	An Aepyceratinae mesophyletid weevil.
Ripidinelia burmiticola[203]	Batelka & Prokop	2019	A ripidiine wedge shaped beetle.
Sanaungulus christensenae[67]	Fanti & Damgaard	2019	A soldier beetle
Sanaungulus curtipennis[68]	Fanti, Damgaard & Ellenberger	2018	A soldier beetle.
Sanaungulus fabriciusi[67]	Fanti & Damgaard	2019	A soldier beetle
Sanaungulus ghitaenoerbyae[68]	Fanti, Damgaard & Ellenberger	2018	A soldier beetle.
Sanaungulus lethi[127]	Fanti & Damgaard	2020	A soldier beetle.
Sanaungulus morellii[127]	Fanti & Damgaard	2020	A soldier beetle.
Sanaungulus rosenzweigi[127]	Fanti & Damgaard	2020	A soldier beetle.
Sanaungulus ruicheni[204]	Hsiao & Huang	2018	A soldier beetle. Originally described as "Ornatomalthinus" ruicheni.[205]
Sanaungulus strungei[67]	Fanti & Damgaard	2019	A soldier beetle
Sanaungulus troelsikloevedali[67]	Fanti & Damgaard	2019	A soldier beetle
Sarothrias cretaceus[206]	Cai et al.	2017	A jacobsoniid beetle.
Scydmaenus linqibini[207]	Yin & Zhou	2020	A scydmaenine ant-like stone beetle.
Scydmaenus minor[208]	Yin & Cai	2019	An ant-like stone beetle.
Scydmobisetia dentipes[209]	Jałoszyński & Bai in Jałoszyński, Bai & Wang	2020	An ant-like stone beetle.
Scydmobisetia loebli[210]	Yin, Zhou & Cai	2018	An ant-like stone beetle.
Scydmobisetia mengjiae[210]	Yin, Zhou & Cai	2018	An ant-like stone beetle.
Scydmobisetia vetutissima[211]	Jałoszyński & Yamamoto in Jałoszyński, Yamamoto & Takahashi	2016	A Glandulariini ant-like stone beetle.
Sieglindea antiqua[81]	Muona	2020	A eucnemine false click beetle.
Sorodites angustipes[212]	Kirejtshuk & Chetverikov	2018	A sap beetle.
Sparedrus archaicus[213]	Vitali & Ellenberger	2019	A false blister beetle
Sphaerothorax uenoi[34]	Cai & Lawrence in Cai et al.	2019	A fringe-winged beetle.
Spinotoma ruicheni[214]	Hsiao & Huang	2018	A Pelecotominae wedge-shaped beetle.
Stegastochlidus saraemcheana[215]	Poinar & Vega	2020	A colydiine ironclad beetle.
Succinumanax birmaniasis[216]	Bao, Rust & Wang	2018	A Eubrianacinae water-penny beetle.
Syntelia sunwukong[217]	Jiang & Wang	2020	A species of Syntelia.
Thescelostrophus cretaceus[218]	Yu et al.	2016	A Eustrophini polypore fungus beetle.
Trihelota fulvata[219]	Tihelka, Huang & Cai	2020	A helotid beetle.
Trixagosoma guangyuani[220]	Li et al.	2020	A throscid beetle.
Vetatractocerus burmiticus[202]	Yamamoto	2019	An atractocerine ship-timber beetle.
Vetatrecus adelfiae[221]	Kypke et al.	2018	An Othiini Staphylininae rove beetle.
Vetatrecus secretum[221]	Kypke et al.	2018	An Othiini Staphylininae rove beetle.
Vetubrachypsectra burmitica[222]	Qu et al.	2020	Abrachypsectrid texas beetle.
Vetudasycerus burmiticus[223]	Cai et al.	2018	A Dasycerinae rove beetle.
Vetujinbrianax cretaceus[224]	Cai & Huang	2018	A water-penny beetle.
Vetuprostomis consimilis[225]	Engel & Grimaldi	2008	A jugular-horned beetle
Vetuproteinus cretaceus[226]	Cai et al.	2016	A Proteininae rove beetle.
Zimmiorhinus conicops[35]	(Clarke & Oberprieler) in Clarke et al.	2018	A Mesophyletinae mesophyletid weevil. Originally described as Elwoodius conicops.[69]

Incertae sedis

Taxon	Authority	Year described	Notes	image
Palaeotylus femoralis[227]	Poinar, Vega, & Legalov	2018	A beetle of uncertain placement.[35]

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