Ommatidae

The Ommatidae are a family of beetles in the suborder Archostemata.[1] The Ommatidae are considered the extant beetle family that has most ancestral characteristics.[2] Extant species of this group only occur in Australia and South America, but the geographical distribution was much wider during the Mesozoic spanning across modern day Europe, Siberia, Myanmar, and China. Discovery of Upper Jurassic Chinese and Upper Cretaceous Burmese amber fossils suggest that they were widespread on Pangea.[3] So far, 25 extinct genera containing over 100 species of these beetles have been described.[4] Three extant genera have been assigned to this family: Omma, Tetraphalerus and Beutelius.[5] The family is considered to be a subfamily of Cupedidae by some authors, but have been found to be more closely related to Micromalthidae by DNA analysis.[6]

Ommatidae
Temporal range: Ladinian–Recent
Omma stanleyi
Tetraphalerus wagneri
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Archostemata
Family: Ommatidae
Sharp & Muir, 1912

Genera

According to Kirejtshuk, 2020.[7]

Extinct genera
  • †Allophalerus Kirejtshuk 2020 Early Jurassic-Late Cretaceous (Asia)
  • †Blapsium Westwood, 1854 Middle Jurassic (Europe)
  • †Brochocoleus Hong, 1982 Early Jurassic-Late Cretaceous (Eurasia)
  • †Bukhkalius Kirejtshuk and Jarzembowski 2020 early Late Cretaceous, (Asia)
  • †Burmocoleus Kirejtshuk 2020 early Late Cretaceous, (Asia)
  • †Cionocoleus Ren, 1995 Middle Jurassic-Early Cretaceous (Eurasia)
  • †Cionocups Kirejtshuk 2020 early Late Cretaceous, (Asia)
  • †Clessidromma Jarzembowski et al. 2017 early Late Cretaceous, (Asia)
  • †Diluticupes Ren 1995 Middle Jurassic-Early Cretaceous (Eurasia)
  • †Echinocups Kirejtshuk and Jarzembowski 2020 early Late Cretaceous, (Asia)
  • †Eurydictyon Ponomarenko, 1969 Early Jurassic (Asia)
  • †Jarzembowskiops Kirejtshuk 2020 early Late Cretaceous, (Asia)
  • †Liassocupes Zeuner 1962 Early Jurassic (Europe)
  • †Lithocupes Ponomarenko, 1966 Late Triassic (Asia)
  • †Miniomma Li, Yamamoto & Cai, 2020, early Late Cretaceous, (Asia)
  • †Monticupes Ren 1995 Middle Jurassic-Early Cretaceous (Asia)
  • †Notocupoides Ponomarenko, 1966 Late Triassic (Asia)
  • †Odontomma Ren et al. 2006 Early Cretaceous (Asia)
  • †Paraodontomma Yamamoto 2017 early Late Cretaceous, (Asia)
  • †Pareuryomma Tan et al. 2012 Early Cretaceous (Asia)
  • †Polyakius Kirejtshuk 2020 early Late Cretaceous, (Asia)
  • †Rhabdocupes Ponomarenko, 1966 Late Triassic (Asia)
  • †Rhopalomma Ashman et al. 2015 Late Jurassic (Australia)
  • †Stegocoleus Jarzembowski and Wang 2016 early Late Cretaceous, (Asia)
  • †Tetraphalerites Crowson 1962 Eocene (Europe)
  • †Zygadenia Handlirsch, 1906 Middle Triassic-Late Cretaceous (Eurasia, ?Australia)

Extant genera
  • Omma Newman, 1839 Late Triassic-Late Cretaceous (Eurasia), recent (Australia)
  • Tetraphalerus Waterhouse, 1901 Middle Jurassic-Early Cretaceous (Asia) recent (South America)
  • Beutelius Escalona et al, 2020 Recent, (Australia)

References
  1. "Australian Faunal Directory". Department of the Environment and Energy. Archived from the original on 2011-03-11. Retrieved 2009-11-21.
  2. Hünefeld, Frank; Marvaldi, Adriana E.; Müller, Bianca; Lawrence, John F.; Beutel, Rolf Georg (2011). "The male postabdomen of the "ancestral" archostematan beetle Tetraphalerus bruchi Heller, 1913 (Ommatidae) and its phylogenetic significance". Arthropod Structure & Development. 40 (2): 146–158. doi:10.1016/j.asd.2010.12.003. PMID 21185393.
  3. Yamamoto, Shûhei (2017). "A new genus of Brochocoleini beetle in Upper Cretaceous Burmese amber (Coleoptera: Archostemata: Ommatidae)". Cretaceous Research. 76: 34–39. doi:10.1016/j.cretres.2017.04.008.
  4. Tan, Jingjing; Wang, Yongjie; Ren, Dong; Yang, Xingke (2012). "New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae". BMC Evolutionary Biology. 12: 113. doi:10.1186/1471-2148-12-113. PMC 3518168. PMID 22776212.
  5. "Ommatidae Species List". Joel Hallan’s Biology Catalog. Texas A&M University. Archived from the original on 2012-10-23.
  6. McKenna, Duane D.; Shin, Seunggwan; Ahrens, Dirk; Balke, Michael; Beza-Beza, Cristian; Clarke, Dave J.; Donath, Alexander; Escalona, Hermes E.; Friedrich, Frank; Letsch, Harald; Liu, Shanlin (2019-12-03). "The evolution and genomic basis of beetle diversity". Proceedings of the National Academy of Sciences. 116 (49): 24729–24737. doi:10.1073/pnas.1909655116. ISSN 0027-8424. PMC 6900523. PMID 31740605.
  7. Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. doi:10.3390/geosciences10020073. ISSN 2076-3263.
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