Indricotheriinae

The subfamily Indricotheriinae are a group of long-limbed, hornless rhinoceroses commonly known as the paraceratheres that evolved in the Eocene epoch and lived until the early Miocene. The first paraceratheres were only about the size of large dogs, growing progressively larger in the late Eocene and Oligocene.[3] They were most common in the rainforest floodplain region which is now Kazakhstan, India, and southwest China, and lived further inland throughout northern and central Asia as well.

Indricotheres Temporal range: Early Eocene to Early Miocene, 54–15 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Reconstructed Paraceratherium skull.
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Perissodactyla
Family:	†Hyracodontidae
Subfamily:	†Indricotheriinae Borissiak, 1923
Genera
†Aralotherium †Buzdartherium †Dzungariotherium †Forstercooperia †Gobioceras †Juxia †Pappaceras †Paraceratherium †Urtinotherium
Synonyms[1]
Baluchitheriinae Osborn, 1923 Paraceratheriinae Osborn, 1923 Forstercooperiidae Kretzoi, 1940[2] Forstercooperiinae Wood, 1963[2]

The paraceratheres reached the peak of their evolution from the middle Oligocene to the early Miocene, where they became very large, herbivorous mammals. Most genera were about the size of modern draft horses and the extinct giant horse Equus giganteus, with some growing significantly larger. The largest genus was Paraceratherium, which was more than twice as heavy as a bull African elephant, and was one of the largest land mammals that ever lived. However, they remained confined to Asia, which at the time was mostly lush lowland floodplains. No fossil remains of paraceratheres have been found in Europe or North America, even though the paraceratheres had millions of years of opportunities to reach those regions. The collision with the Indian subcontinent and the Himalayan uplift led to global cooling, desertification, and the disappearance of forest habitats, which resulted in the extinction of these giant ungulates.

Although considered a subfamily of the Hyracodontidae by some authors, recent authors treat the paraceratheres as a distinct family, Paraceratheriidae (Wang et al. 2016 recover hyracodonts as more basal than paraceratheres).[4][5]