Araneomorphae

The Araneomorphae (also called the Labidognatha) are an infraorder of spiders. They are distinguished by having chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their close kin), where they point straight down. Most of the spiders that people encounter in daily life belong to the Araneomorphae.

Araneomorph spiders
Temporal range: Triassic–present
Nephila inaurata (Nephilidae)
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Suborder: Opisthothelae
Infraorder: Araneomorphae
Families

see Table of families

Diversity
95 families
Hippasa agelenoides  common funnel web spider

Distinguishing characteristics

Note the difference in the orientations of the chelicerae (fangs) of the two spiders below, representatives of the Mygalomorphae and the Araneomorphae. The number of book lungs (when present) can also help distinguish between members of these two major groups.

Mygalomorphae

This Atrax robustus (a member of the Hexathelidae) is presenting a defensive posture, and by so doing clearly shows the orientation of its chelicerae, which go up and down and parallel to the long axis of the spider's body, as with other representatives of the Mygalomorphae.

Araneomorphae

In the Araneomorphae, the fangs slope towards each other, giving these spiders many more possibilities than the Mygalomorphae, which can only bite top down. In contrast to the Mygalomorphae, where females can live for many years, most Araneomorphae die after about a year.[1]

Spiders included

Almost all of the familiar spiders are included in this group. The major exception is the Tarantulas, which have become so common as pets that many people have seen them. There are a few other members of Mygalomorphae that one might see around homes or gardens, but they typically are relatively small and not easily noticed. For instance, the females of one such species lives and hunts from within a long silken tube, so unless one opens the tube or chances upon a male looking for a mate, one will never see them. The Araneomorphae, to the contrary, include the weavers of spiral webs, the cobweb spiders that live in the corners of our rooms and between windows and screens, the crab spiders that lurk on the surfaces of the flowers in our gardens, the jumping spiders that look back at us curiously from walls and tree trunks, the wolf spiders that sometimes carpet good hunting sites in a sunny spot in the lawn, the large Huntsman spiders that sometimes frighten people by getting into their cars or taking up residence behind wall clocks.

Systematics

In older schemes, the Araneomorphae are divided into two lineages, the Hypochilae (containing only the family Hypochilidae), and the Neocribellatae. The Neocribellatae are in turn divided into the Austrochiloidea, and the two series Entelogynae and Haplogynae, each containing several superfamilies:

A cladogram shows the relation among taxa:[2]

   Opisthothelae   

Mygalomorphae

   Araneomorphae   
   

Hypochilidae

   Austrochiloidea   
   

Gradungulidae

Austrochilidae

   Araneoclada   

Haplogynae

Entelegynae

Most spiders in the Haplogynae series have six eyes, while most of those in the Entelegynae series have eight.

Table of families

Key
Genera1≥2≥10≥100
Species1–9≥10≥100≥1000
Araneomorphae families[note 1]
FamilyGeneraSpeciesCommon nameExample
Agelenidae781282araneomorph funnel-web spidersHobo spider
Amaurobiidae49274tangled nest spidersCallobius claustrarius
Ammoxenidae418
Anapidae58223
Anyphaenidae56563anyphaenid sac spidersYellow ghost spider
Araneidae1743128orb-weaver spidersZygiella x-notata
Archaeidae590pelican spidersMadagascarchaea gracilicollis
Arkyidae237
Austrochilidae310Tasmanian cave spider
Caponiidae18119Diploglena capensis
Cheiracanthiidae
(syn. Eutichuridae)
12351Cheiracanthium mildei
Cithaeronidae28
Clubionidae15618sac spidersClubiona trivialis
Corinnidae67779dark sac spidersCastianeira sp.
Ctenidae47525tropical wolf spidersBrazilian wandering spiders
Cyatholipidae2358
Cybaeidae19259
Cycloctenidae880
Deinopidae265net-casting spidersRufous net-casting spider
Desidae60297intertidal spidersPhryganoporus candidus
Dictynidae52464Nigma walckenaeri
Diguetidae215coneweb spiders
Drymusidae217false violin spiders
Dysderidae24564woodlouse hunter spidersWoodlouse spider
Eresidae998velvet spidersEresus sandaliatus
Filistatidae19164crevice weaversSouthern house spider
Gallieniellidae1056
Gnaphosidae[note 2]1582532flat-bellied ground spidersDrassodes cupreus
Gradungulidae716large-clawed spidersCarrai cave spider
Hahniidae23346dwarf sheet spiders
Hersiliidae16181tree trunk spidersHersilia savignyi
Homalonychidae13
Huttoniidae11Huttonia palpimanoides
Hypochilidae212lampshade spidersHypochilus thorelli
Lamponidae23192White-tailed spider
Leptonetidae21346Tooth cave spider
Linyphiidae6074566dwarf / money spidersLinyphia triangularis
Liocranidae31272liocranid sac spiders
Lycosidae1242419wolf spidersLycosa tarantula
Malkaridae1146shield spiders
Mecysmaucheniidae725
Megadictynidae22
Mimetidae12152pirate spidersOarces reticulatus
Miturgidae29130long-legged sac spiders
Mysmenidae13137spurred orb-weavers
Nesticidae16278cave cobweb spidersNesticella marapu
Nicodamidae727
Ochyroceratidae20216midget ground weaversTheotima minutissima
Oecobiidae6113disc web spidersOecobius navus
Oonopidae1141801dwarf hunting spidersOonops domesticus
Orsolobidae30188
Oxyopidae9457lynx spidersGreen lynx spider
Pacullidae438
Palpimanidae18150palp-footed spiders
Penestomidae19
Periegopidae13
Philodromidae30539philodromid crab spidersPhilodromus dispar
Pholcidae771666daddy long-legs spidersPholcus phalangioides
Phrurolithidae13205
Physoglenidae1372
Phyxelididae1464
Pimoidae441Pimoa cthulhu
Pisauridae51356nursery web spidersPisaura mirabilis
Plectreuridae231
Psechridae261
Salticidae6356080jumping spidersZebra spider
Scytodidae5248spitting spidersScytodes thoracica
Segestriidae4130tubeweb spidersSegestria florentina
Selenopidae10257wall spidersSelenops radiatus
Senoculidae131
Sicariidae31623recluse spidersBrown recluse
Sparassidae881224huntsman spidersAvondale spider
Stenochilidae213
Stiphidiidae20125Tartarus mullamullangensis
Symphytognathidae873dwarf orb-weaversPatu digua
Synaphridae313
Synotaxidae111
Telemidae1079long-legged cave spiders
Tetrablemmidae27129armored spiders
Tetragnathidae48996long jawed orb-weaversOrchard spider
Theridiidae1242503cobweb spidersRedback spider
Theridiosomatidae19124ray spidersTheridiosoma gemmosum
Thomisidae1702171crab spidersGoldenrod spider
Titanoecidae553Goeldia obscura
Toxopidae1482
Trachelidae18232
Trechaleidae16120
Trochanteriidae19153
Trogloraptoridae11Trogloraptor marchingtoni
Udubidae415
Uloboridae19283hackled orb-weaversUloborus walckenaerius
Viridasiidae29
Xenoctenidae433
Zodariidae851141ant spidersZodarion germanicum
Zoropsidae26180Zoropsis spinimana

References

Notes

  1. Unless otherwise shown, currently accepted families and counts based on the World Spider Catalog version 19.0 as of 11 July 2018.[3] In the World Spider Catalog, "species" counts include subspecies. Assignment to sub- and infraorders based on Coddington (2005, p. 20) (when given there).
  2. June 2019 data
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