Itaboraian

The Itaboraian (Portuguese: Itaboraiense) age is a period within the Early Eocene geologic time (53.0–50.0 Ma) epoch of the Paleogene, used more specifically with South American land mammal ages (SALMA). It follows the Riochican and precedes the Casamayoran age.[1]

Stratigraphy and location of the Itaboraí Basin.
Crocodile skull indicates Sahitisuchus find.

Etymology

This age is named after the Itaboraí Formation in the Itaboraí Basin in Itaboraí, a Greater Rio municipality in the state of Rio de Janeiro in Brazil not very far away from Niterói.

Formations
Locations of Itaboraian formations
Formation
bold is type		 Country Basin Notes
Itaboraí Formation BrazilItaboraí Basin
[2]
Bogotá Formation ColombiaAltiplano Cundiboyacense
[3]
Cerrejón Formation ColombiaCesar-Ranchería Basin
[4]
Chota Formation PeruBagua Basin
[5][6]
Koluel Kaike Formation ArgentinaGolfo San Jorge Basin
[7]
Las Flores Formation ArgentinaGolfo San Jorge Basin
[8]
Maíz Gordo Formation ArgentinaSalta Basin
[9]
Mogollón Formation PeruTalara & Tumbes Basins
[10]
Muñani Formation PeruAltiplano Basin
[11]

Fossils
Group Fossils Formation Notes
MammalsAsmithwoodwardia scotti, Austropediomys marshalli, Bergqvistherium primigenia, Bobbschaefferia fluminensis, Camargomendesia pristina, Carodnia vieirai, Carolocoutoia ferigoloi, Carolopaulacoutoia itaboraiensis, Colbertia magellanica, Derorhynchus singularis, Didelphopsis cabrerai, Eobrasilia coutoi, Epidolops ameghinoi, Gashternia carioca, Gaylordia macrocynodonta, Gaylordia mater, Guggenheimia brasiliensis, Guggenheimia crocheti, Homalostylops atavus, Itaboraidelphys camposi, Lamegoia conodonta, Marmosopsis juradoi, Miguelsoria parayirunhor, Minusculodelphis minimus, M. modicum, Mirandatherium alipioi, Monodelphopsis travassosi, Palaeocladosictis mosesi, Paranisolambda prodromus, Patene simpsoni, Periprotodidelphis bergqvistae, Procaroloameghinia pricei, Protodidelphis mastodontoides, P. vanzolinii, Protolipterna ellipsodontoides, Ricardocifellia protocenica, Riolestes capricornicus, Riostegotherium yanei, Tetragonostylops apthomasi, Victorlemoinea prototypica, Xenodelphis doelloi, Zeusdelphys complicatus, ?Arminiheringia sp., Henricosbornia sp., Australidelphia indet., Borhyaenidae indet., Didelphidae indet., Didolodontidae indet., Herpetotheriidae indet., Microbiotheriidae indet., Notoungulata indet., Paucituberculata indet., Pediomyidae indet., Peradectidae indet., Polydolopidae indet., Protodidelphidae indet.Itaboraí
[12]
[13][14]
[15]
[2][16]
[17]
Ernestokokenia yirunhor, Gashternia ctalehor, Henricosbornia waitehor, Seudenius cteronc, Shecenia ctirneru, ?Isotemnus ctalego, ?Peripantostylops orehor, ?Polydolops kamektsen, Kibenikhoria sp., Archaeopithecidae indet., Borhyaenidae indet., ?Litopterna indet., ?Notostylopidae indet.Koluel Kaike
[18]
Anisolambda sp., Antepithecus sp., Asmithwoodwardia sp., Bardalestes sp., Bobbschaefferia sp., Camargomendesia sp., Carolopaulacoutoia sp., Colbertia sp., Derorhynchus sp., Didelphopsis sp., Epidolops sp., Ernestokokenia sp., Gashternia sp., Guggenheimia crocheti, Henricosbornia sp., Isotemnus sp., Itaboraitherium sp., Kibenikhoria sp., Marmosopsis sp., Mirandatherium sp., Monodelphopsis sp., Peripantostylops sp., Polydolops sp., Procaroloameghinia pricei, Protodidelphis sp., Shecenia sp., Trigonostylops sp., Victorlemoinea sp., Woodburnodon sp., Ceratodontidae indet., ?Patene indet.Las Flores
[19]
Carodnia inexpectansMogollón
[20][21]
Chulpasia mattaueri, Peradectes austrinum, Perutherium altiplanense, Sillustania quechuense, Umayodus raimondi, Didelphidae indet., ?Didolodontidae indet., Notoungulata sp., Pediomyidae indet., Proteutheria indet.Muñani
[22]
[23][24]
BirdsDiogenornis fragilis, Eutreptodactylus itaboraiensis, Itaboravis elaphrocnemoides, Paleopsilopterus itaboraiensisItaboraí
[16][25]
[26][27]
Latidae indet., Siluriformes indet., Tetragonopterinae indet.Muñani
[24]
Reptiles &
amphibians		Acherontisuchus guajiraensis, Anthracosuchus balrogus, Carbonemys cofrinii, Cerrejonemys wayuunaiki, Cerrejonisuchus improcerus, Pelomedusoides sp., Puentemys mushaisaensis, Titanoboa cerrejonensisCerrejón
[28][29]
?Madtsoia sp.Koluel Kaike
[18]
Notocaiman stromeriPeñas
Coloradas
[30]
Bretesuchus bonaparteiMaíz Gordo
[31]
Apodops pricei, Coniophis cf. precedens, Corallus priscus, Hechtophis austrinus, Hoffstetterella brasiliensis, Itaboraiophis depressus, Madtsoia camposi, Paraungaliophis pricei, Paulacoutophis perplexus, Sahitisuchus fluminensis, Waincophis cameratus, Waincophis pressulus, Xenopus romeri, ? Russellophiidae indet.Itaboraí
[32][33]
Lorosuchus nodosus, Pelomedusoides cf. argentinensisRío Loro
[34]
cf. Coniophis sp., ?Roxochelys vilavilensis, Crocodylia indet., ?Leptodactylidae indet.Muñani
[24]
FishesDipnoi sp., Elopomorpha sp.Cerrejón
[28]
Lepidosiren cf. paradoxa, Ceratodus sp.Muñani
[24]
MolluscsBulimulus fazendicusItaboraí
[35]
FloraMenispina evidensBogotá
[36]
Stephania palaeosudamericanaBogotá, Cerrejón
[37]
Malvaciphyllum macondicus, Petrocardium cerrejonense, Petrocardium wayuuorumCerrejón
[38][39]

Correlations
Itaboraian correlations in South America
FormationItaboraíLas FloresKoluel KaikeMaíz GordoMuñaniMogollónBogotáCerrejónYpresian (IUCS) • Wasatchian (NALMA)
Bumbanian (ALMA) • Mangaorapan (NZ)
BasinItaboraíGolfo San JorgeSaltaAltiplano BasinTalara &
Tumbes		Altiplano
Cundiboyacense		Cesar-Ranchería
Itaboraian (South America)
Country Brazil Argentina Peru Colombia
Carodnia
Gashternia
Henricosbornia
Victorlemoinea
Polydolopimorphia
Birds
Reptiles
Fish
Flora
EnvironmentsAlluvial-lacustrineAlluvial-fluvialFluvio-lacustrineLacustrineFluvialFluvio-deltaic
Itaboraian volcanoclastics

Itaboraian fauna

Itaboraian flora
VolcanicYes

References
  1. Paleo Database: Itaboraian
  2. Oliveira & Goin, 2011
  3. Bayona et al., 2012, p.103
  4. Jaramillo et al., 2007, p.168
  5. Chacaltana et al., 2015, p.2
  6. Castro Medina, 2010, p.53
  7. Raigemborn et al., 2010, p.251
  8. Raigemborn et al., 2010, p.249
  9. Narváez, 2009, p.24
  10. Quispe Alcalá, 2010, p.21
  11. Gelfo & Goin, 2011
  12. Oliveira et al., 2016
  13. Carneiro et al., 2018
  14. Carneiro, 2018
  15. De Oliveira & Goin, 2015
  16. São José de Itaboraí at Fossilworks.org
  17. São José 700 m at Fossilworks.org
  18. Cañadón Hondo at Fossilworks.org
  19. Las Flores at Fossilworks.org
  20. Antoine et al., 2015
  21. Carodnia inexpectans at Fossilworks.org
  22. Gelfo & Sigé, 2011
  23. Laguna Umayo at Fossilworks.org
  24. LURMU at Fossilworks.org
  25. Diagenornis at Fossilworks.org
  26. Itaboravis at Fossilworks.org
  27. Paleopsilopterus at Fossilworks.org
  28. La Puente Pit at Fossilworks.org
  29. Anthracosuchus at Fossilworks.org
  30. Notocaiman stromeri at Fossilworks.org
  31. Bretesuchus at Fossilworks.org
  32. Itaboraí snakes at Fossilworks.org
  33. Portland Quarry at Fossilworks.org
  34. El Cadillal at Fossilworks.org
  35. Salvador & Simone, 2013, p.7
  36. Menispina evidens at Fossilworks.org
  37. Stephania palaeosudamericana at Fossilworks.org
  38. Cerrejón 0315 at Fossilworks.org
  39. Coalbeds 100 at Fossilworks.org

Bibliography
Itaboraí Formation
Bogotá Formation
Cerrejón Formation
Chota Formation
Las Flores Formation
Koluel Kaike Formation
Maíz Gordo Formation
Mogollón Formation
Muñani Formation
Peñas Coloradas Formation
Río Loro Formation
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