Huayquerian

The Huayquerian (Spanish: Huayqueriense) age is a period of geologic time (9.0–6.8 Ma) within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification. It follows the Mayoan and precedes the Montehermosan age.[1]

Etymology

The age is named after the Huayquerías Formation in the western Cuyo Basin of northwestern Argentina that was later dated to the Montehermosan. The most complete Huayquerian fauna is found in the Cerro Azul Formation, in Buenos Aires Province also referred to as Epecuén Formation.

Formations
Locations of Huayquerian formations
Formation
bold is type		 Country Basin Notes
Huayquerías Formation (now Montehermosan) ArgentinaCuyo Basin
[2][3]
Andalhuala Formation ArgentinaSanta María-Hualfín Basin
[4]
Bahía Inglesa Formation - Cerro Ballena ChileCaldera Basin
[5]
Camacho Formation UruguayNorte & Santa Lucía Basins
[6]
Cerro Azul Formation ArgentinaColorado Basin
[7][8][9]
Chiquimil Formation ArgentinaSanta María-Hualfín Basin
[4][10]
Coquimbo Formation ChileTongoy Bay
[11]
Las Flores Formation ArgentinaSierra del Tontal
[12]
Entrerriana Formation ArgentinaChaco Basin
[note 1]
Epecuén Formation ArgentinaClaromecó Basin
[note 2]
Içá Formation BrazilMadre de Deus Basin
[13]
Iñapari Formation PeruMadre de Dios Basin
[14]
India Muerta Formation ArgentinaSierras Pampeanas
[15]
Ituzaingó Formation ArgentinaParaná Basin
[16][17]
Kiyu Formation UruguaySanta Lucía Basin
[18][note 3]
Madre de Dios Formation PeruMadre de Dios Basin
[20]
Maimará Formation ArgentinaEastern Argentinian Andes
[21]
Mauri Formation BoliviaAltiplano Basin
[22]
Miramar Formation PeruSechura Basin
[23]
El Morterito Formation ArgentinaCajón Valley
[24]
Muyu Huasi Formation BoliviaMuyu Huasi Basin
[25]
Navidad Formation ChileChilean Coast Range
[26]
Palo Pintado Formation ArgentinaSalta Basin
[27]
Paraná Formation ArgentinaChaco Basin
[17][28][29]
Pebas Formation Brazil
 Colombia
 Ecuador
 Peru		Amazon Basin
[30][31]
Piquete Formation ArgentinaSalta Basin
[32]
Pisco Formation PeruPisco Basin
[33][34]
Quehua Formation BoliviaAltiplano Basin
[35]
Raigón Formation UruguayNorte Basin
[36]
Rosa Pata Formation BoliviaAltiplano Basin
[37]
Saldungaray Formation ArgentinaBuenos Aires Province
[9]
Salicas Formation ArgentinaSierra de Velasco
[38]
Solimões Formation BrazilSolimões Basin
[39]
Urumaco Formation VenezuelaFalcón Basin
[40]

Fossil content
Group Fossils Formation Notes
MammalsCyonasua pascuali, Huayqueriana cristataHuayquerías
[41][42]
Australophoca changorum, Brachydelphis jahuayensis, Thalassocnus natans, Acrophoca sp., Balaenoptera sp., ?Nanosiren sp., Odobenocetops sp., Piscophoca sp., Pliopontos sp., Pontistes sp., Balaenopteridae indet., Delphinoidea indet., Istiophoridae indet., Kogiidae indet., Monachinae indet., Odontoceti indet., Phocidae indet., Phocoenidae indet., Physeteroidea indet., Xiphiidae indet.Bahía
Inglesa
[43][44]
[45][46]
Arazamys castiglionii, Kiyumylodon lecuonai, Kraglievichia paranense, Lestobradys sprechmanni, Neoglyptatelus uruguayensis, Pseudoplohophorus absolutus, Euphractini indet.Camacho
[47][48]
[49][50]
[51]
Hyperdidelphys pattersoni, Microtragulus rusconii, Neocavia pampeana, Palaeoctodon aff. simplicidens, Pampamys emmonsae, Pliolestes venetus, Reigechimys plesiodon, R. simplex, Thylacosmilus atrox, Thylamys pinei, Thylatheridium dolgopolae, T. hudsoni, Zygolestes tatei, Eumysops sp., Sparassocynidae indet.Cerro
Azul
[52][53]
[54][55]
[56][57]
Eosclerocalyptus planus, Orthomyctera andina, ?Palaeocavia mawka, Cardiomys sp., Chasicotatus sp., Chorobates sp., Diadiaphorus sp., Gyriabrus sp., Lagostomus (Lagostomopsis) sp., Paedotherium sp., Paleuphractus sp., Paranamys sp., Potamarchus sp., Proscelidodon sp., Protabrocoma sp., Stromaphorus sp., Tetrastylus sp., Typotheriopsis sp., Vassallia sp., Vetelia sp., Xotodon sp.Chiquimil
[58][59]
Balaena dubia, Notictis ortizi, Ischyrorhynchus vanbenedeni, Otaria fischeri, Palaeobalaena bergi, P. dubia, Phugatherium cataclisticum, Pontistes rectifrons, Pontivaga fischeri, Proterotherium americanum, Ribodon limbatus, Saurocetes argentinus, Scalabrinitherium paranense, Stylocynus paranensis, Toxodon paranensis, Chironectes sp.Entrerriana
[60][61]
Aspidocalyptus castroi, Borhyaenidium musteloides, Chorobates villosissimus, Coscinocercus brachyurus, C. marcalaini, Cyonasua brevirostris, C. prebrevirostris, Doellotatus chapadmalensis, Elassotherium altirostre, Eoauchenia cingulata, Epecuenia thoatherioides, Hemihegetotherium lazai, Macrauchenia latidens, Macrochorobates scalabrinii, Macroeuphractus morenoi, Paedotherium minor, Phtoramys hidalguense, Pisanodon nazari, Plohophorus araucanus, Pseudotypotherium carhuense, Tetrastylus araucanus, Thylacosmilus atrox, Thylatheridium dolgopolae, T. hudsoni, Cardiomys sp., Eosclerocalyptus sp., Lagostomus sp., Lutreolina sp., Orthomyctera sp., Palaeocavia sp., Paleuphractus sp., Plesiomegatherium sp., Proeuphractus sp., Promacrauchenia sp., Thylamys sp., Hydrochoeridae indet.Epecuén
[62]
Cardiatherium calingastaenseLas Flores
[12]
Amazonycteris divisusIçá
[13]
Mionothropus cartelleiIñapari
[63]
"Microtragulus ameghino"India
Muerta
[15]
?Adinotherium paranense, Amphiocnus paranense, Brachytherium cuspidatum, Cardiatherium paranense, Chasicotatus spinozai, Chlamyphractus pressulus, Coelosoma eversa, Comaphorus concisus, Cyonasua argentina, Dasypus neogaeus, Diadiaphorus paranensis, Dilobodon lutarius, Dinotoxodon paranensis, Eleutherocercus paranensis, Eomegatherium nanum, Eumysops parodii, Eutomodus elautus, Haplodontherium limun, Hoplophorus verus, Kraglievichia paranense, Licaphrium mesopotamiense, Macroeuphractus retusus, Megabradys darwini, Megalonychops primigenius, Mesopotamocnus brevirostrum, Munizia paranensis, Neobrachytherium ameghinoi, Neoepiblema ambrosettianus, N. horridula, Neohapalops rothi, Notictis ortizi, Octomylodon aversus, Orthotherium laticurvatum, O. robustum, Oxyodontherium piramydatus, O. zeballosi, Pachynodon modicus, Palaehoplophorus antiquus, Palaeotoxodon paranensis, ?P. protoburmeisteri, ?P. virgatus, Paraglyptodon paranensis, Parahoplophorus paranensis, Paranabradys vucetichae, Paranauchenia denticulata, Philander entrerianus, Phoberomys burmeisteri, P. insolita, P. lozanoi, P. minima, P. praecursor, Phugatherium cataclisticum, Pliomegatherium lelongi, Pliomorphus mutilatus, P. paranensis, P. robustus, Proeuphractus limpidus, Prolestodon antiquus, P. paranensis, Promacrauchenia antiqua, Promegatherium parvulum, P. smaltatum, Promylodon paranensis, Pronothrotherium mirabilis, Protoglyptodon primiformis, Protomegalonyx doellojuradoi, P. praecursor, Proterotherium cervioides, P. gradatum, Protypotherium antiquum, Pseudoeuryurus lelongianus, Ranculcus scalabrinianus, Saurocetes gigas, Scalabrinitherium bravardi, S. rothii, Scirrotherium carinatum, Sphenotherus paranensis, Stenotephanos plicidens, Strabosodon acuticavus, S. obtusicavus, Stylocynus paranensis, Thylacosmilus atrox, Torcellia paranense, Toxodontherium compressum, T. reverendum, ?Trachycalyptus cingulatus, Urotherium interundatum, Xotodon doellojuradi, Xotodon foricurvatus, Zygolestes paranensis, Anatochoerus sp., ?Berroia sp., Berthawyleria sp., Briaromys sp., Cardiomys sp., Carlesia sp., Caviodon sp., Chironectes sp., Colpostemma sp., Contracavia sp., Cullinia sp., Diaphoromys sp., Doellomys sp., Drytomomys sp., Eumegamys sp., Eumegamysops sp., Gyriabrus sp., Haplostropha sp., Isostylomys sp., Lagostomus (Lagostomopsis) sp., Myocastor sp., Palaeocavia sp., Paradoxomys sp., Paranamys sp., Parodimys sp., Pentastylomys sp., Pliodolichotis sp., Potamarchus sp., Prodolichotis sp., Protabrocoma sp., Protomegamys sp., Pseudosigmomys sp., Pyramiodontherium sp., Strophostephanos sp., Tetrastylus sp., Trachytypotherium sp., ?Zaedyus sp., Iniidae indet., Scelidotheriinae indet.Ituzaingó
[64]
[65][66]
[67]
Proeuphractus limpidusKiyu
[68]
[note 4]
Noctilio lacrimaelunaris, Surameryx acrensis, Sylvochoerus woodburnei, Waldochoerus bassleriMadre de
Dios
[69][70]
[71][72]
[73]
Proscelidodon patrius, Sparassocynus maimarai, Xotodon maimarensisMaimará
[74][75]
[76]
Borhyaenidium altiplanicus, Hoffstetterius imperator, Trachycalyptoides achirense, Xyophorus villarroeli, Dasypodidae indet., Hegetotheriidae indet., Lagostominae indet., Mesotheriidae indet., Nothrotheriinae indet., Sclerocalyptinae indet., Toxodontidae indet.Mauri
[77]
Balaenopteridae indet.Miramar
[78]
Calchaquitherium mixtumEl Morterito
[79]
?Cardiomys sp., cf. Doellotatus sp., Lagostomus (Lagostomopsis) sp., ?Orthomyctera sp., cf. Paleuphractus sp., Protabrocoma sp., Protypotherium sp., Pseudohegetotherium sp., cf. Sciamys sp., Dinomyidae indet., Erethizontidae indet., Mylodontidae indet., Proterotheriidae indet.Muyu
Huasi
[80]
Paedotherium kakaiPalo
Pintado
[81]
Dioplotherium sp., Metaxytherium sp.Paraná
[82][83]
Koristocetus pescei, Tiucetus rosaePisco
[84][85]
Cranithlastus xibiensis, Megatheriinae indet.Piquete
[86]
Microtypotherium cf. choquecotense, Plesiotypotherium cf. achirense, P. cf. majus, P. minus, Orthomyctera sp., Dasypodidae indet.Quehua
[87]
Pronothrotherium figueirasi, P. mirabilisRaigón
[88]
Pseudotypotherium sp., Dasypodidae indet., Glyptodontidae indet., ?Macraucheniidae indet.Rosa Pata
[89]
Chasicotatus ameghinoi, C. cf. peiranoi, Chorobates villosissimus, Doellotatus cf. inornatus, D. cf. praecursor, Macroeuphractus cf. morenoi, Neophanomys biplicatus, Paedotherium bonaerense, P. cf. minor, Phtoramys cf. hidalguense, P. homogenidens, Ringueletia simpsoni, Tremacyllus cf. impressus, Xenodontomys ellipticus, Acantharodeia sp., Aspidocalyptus sp., Berthawyleria sp., cf. Borhyaenidium sp., Dolicavia sp., Lagostomus sp., Neocavia sp., Palaeocavia sp., Promacrauchenia sp., Caviidae indet., Dasypodidae indet.Saldungaray
[90][91]
[92][93]
[94]
Eosclerocalyptus planus, Hemihegetotherium cf. torresi, Lagostomus cf. pretrichodactyla, Macrochorobates scalabrinii, Neophanomys biplicatus, Orthomyctera andina, Paedotherium minor, Proeuphractus limpidus, Acantharodeia sp., cf. Cardiomys sp., Chaetophractus sp., Chasicotatus sp., Lagostomus sp., Neobrachytherium sp., Potamarchus sp., Protypotherium sp., Pseudotypotherium sp., cf. Tremacyllus sp., Hoplophorini indet., Octodontidae indet.Salicas
[95][96]
Abothrodon pricei, Acrecebus fraileyi, Amahuacatherium peruvium, Didelphis solimoensis, Eumegamys paranensis, Ferigolomys pacarana, Gyrinodon quasus, Ischyrorhynchus vanbenedeni, Lutreolina materdei, Mesenodon juruaensis, Mesotoxodon pricei, Minitoxodon acrensis, Neoepiblema ambrosettianus, N. horridula, Neotoxodon pascuali, Neotrigodon utoquineae, Noctilio lacrimaelunaris, Octodontobradys puruensis, Phoberomys bordasi, P. burmeisteri, P. minima, Plesiotoxodon amazonensis, Plicodontinia mourai, cf. Pomatodelphis bobengi, Potamarchus adamiae, P. murinus, P. sigmodon, Pseudoprepotherium venezuelanum, Purperia cribatidens, Ribodon limbatus, Scirrotherium carinatum, Scleromys colombianus, Solimoea acrensis, Stenodon campbelli, Telicomys amazonensis, Toxodontherium listai, Trigodonops lopesi, Urumacotherium campbelli, Xenastrapotherium amazonense, Anadasypus sp., Asterostemma sp., Cardiatherium sp., Cullinia sp., Gyriabrus sp., cf. Hapalops sp., Kraglievichia sp., Neoglyptatelus sp., Palaeotoxodon sp., Paraglyptodon sp., Paratrigodon sp., cf. Planops sp., Pliomorphus sp., Plohophorus sp., cf. Protomegalonyx sp., cf. Proterotherium sp., Pseudoprepotherium sp., Ranculcus sp., Saurocetes sp., Simplimus sp., Stirtonia sp., Tetrastylus sp., Trigodon sp., Urumacotherium sp., Cardiomyinae indet., Dasyproctidae indet., Erethizontidae indet., Heteropsomyinae indet., Macraucheniidae indet., Marsupialia indet., Megalonychidae indet., Molossidae indet., Mylodontidae indet., Neoepiblemidae indet., Pampatheriinae indet., Pontoporiidae indet., Toxodontidae indet.Solimões
[97][98]
[99][100]
[101][102]
[103]
Ullumys pattoniLas Tapias
[104]
Eionaletherium tanycnemius, Lestodon urumaquensis, Phoberomys pattersoni, Pattersonocnus diazgameroi, Urumacocnus urbanii, Cardiatherium sp., Eumegamys sp., Hemihegetotherium sp., Ocnerotherium sp., Ribodon sp., Tetrastylus sp.Urumaco
[105][106]
[107][108]
[109][110]
[111]
BirdsArgentavis magnificensAndalhualá
[112]
Pelagornis chilensis, Spheniscidae indet.Bahía Inglesa
[45][113]
Argentavis magnificens, Eudromia sp.Epecuén
[62]
Andalgalornis sp., Devincenzia sp., Megapaloelodus sp., Phoenicopteridae indet., Phorusrhacidae indet., Rallidae indet., Rheidae indet.Ituzaingó
[65]
Anhinga fraileyi, A. cf. grandis, A. minuta, Phorusrhacinae indet.Solimões
[99][100]
ReptilesCaiman australis, C. paranensis, C. praecursor, Leptorramphus entrerrianus, Rhamphostomopsis neogaeus, Testudines indet.Entrerriana
[60]
Caiman australis, C. gasparinae, C. jacare, C. latirostris, C. lutescens, C. cf. yacare, Gryposuchus neogaeus, Mourasuchus arendsi, Parahydraspis paranaensis, Phrynops cf. geoffroanus, Tupinambis cf. merianaeItuzaingó
[65][66]
Chelonioidea indet., Crocodylia indet., Eusuchia indet.Miramar
[78]
Caiman cf. latirostrisPalo
Pintado
[114]
Caiman brevirostris, C. niteroensis, Charactosuchus fieldsi, C. mendesi, C. sansoai, Chelus colombianus, Chelus lewisi, Colombophis portai, C. spinosus, Gryposuchus jessei, Mourasuchus amazonensis, M. arendsi, Podocnemis bassieri, Podocnemis negrii, Purussaurus brasiliensis, Chelonoidis sp., aff. Epicrates sp., Eunectes sp., Hesperogavialis sp., cf. Paradracaena sp., Stupendemys sp., Waincophis sp., ?Brachygnathosuchus sp., Colubridae indet., ?Gavialosuchus sp., Pelomedusidae indet., Serpentes indet., Testudinidae indet.Solimões
[99][100]
[101][115]
Melanosuchus fisheri, Mourasuchus pattersoni, Stupendemys geographicusUrumaco
[116][117]
FishesCarcharodon carcharias, C. hastalis, Cosmopolitodus hastalisBahía Inglesa
[43][44]
Megalodon, Carcharhinus egertoni, Carcharodon hastalis, Carcharias sp., Myliobatis sp.Camacho
[118][119]
[120][121]
Lamna amplibasidens, L. serridens, L. unicuspidens, Myliobatis americanus, Odontaspis macrota, Osteophorus typus, Sargus incertus, Silurus agassizii, Sparus antiquus, Squalus eocenus, Squalus obliquidens, Solea sp.Entrerriana
[60]
Colossoma macropomus, Megapiranha paranensis, Cynodontidae indet.Ituzaingó
[65][66]
Megalodon, Carcharhinus brachyurus, Carcharhinus falciformis, Carcharhinus leucas, Carcharias taurus, Galeocerdo aduncus, Ginglymostoma cirratum, Hemipristis serra, Negaprion brevirostris, Sphyrna lewini, Sphyrna zygaena, Aetobatus sp., Dasyatis sp., Myliobatis sp.Miramar
[78]
Megalodon, Carcharias taurus, Carcharhinus sp., Carcharodon plicatilis, Galeocerdo aduncus, Hemipristis serra, Megascyliorhinus trelewensis, Dasyatis sp., Heterodontus sp., Sphyrna sp., Squatina sp., Ariidae indet., Holocephali indet., Rajidae indet., Sciaenidae indet., Sparidae indet.Paraná
[122]
Phractocephalus acreornatus, Colossoma sp., Callichthyidae indet., Doradidae indet., Osteoglossidae indet., Pimelodidae indet., Potamotrygonidae indet.Solimões
[99][100]
DecapodsHomarus meridionalisEntrerriana
[60]
FloraAlnipollenites verus, Annulispora folliculosa, Arecipites asymmetricus, Baculatisporites comaumensis, Biretisporites potoniaei, Chenopodipollis chenopodiaceoides, C. multicavus, Cricotriporites guianensis, Cyathidites congoensis, C. minor, Cyperaceaepollis neogenicus, Deltoidospora minor, Dictyophyllidites arcuatus, D. chiquimilense, D. mortoni, Echiperiporites parviechinatus, Equisetosporites claricristatus, E. lusaticus, E. notensis, Foveotriletes microfoveolatus, Gleicheniidites senonicus, Graminidites media, Haloragacidites trioratus, Laevigatosporites ovatus, Leiotriletes regularis, Leptolepidites major, Liliacidites vermireticulatus, Malvacipollis argentina, Muricingulisporis verrucosus, Nothofagidites saraensis, Osmundacidites ciliatus, Palaeocoprosmadites zelandiae, Parsonsidites multiporus, Periporopollenites vesicus, Plicifera delicata, Podocarpidites ellipticus, P. exiguus, P. marwickii, Polycolporopollenites esobalteus, Polypodiaceoisporites simplex, P. tumulatus, Polypodiisporites speciosus, Psilamonocolpites grandis, Psilaperiporites minimus, Psilatricolporites cyamus, Reticuloidosporites tenellis, Rhoipites alveolatus, R. baculatus, R. cienaguensis, R. exiguus, R. karamuensis, R. santafesii, Rugulatisporites caperatus, Sparganiaceaepollenites delicata, Stereisporites antiquasporites, Striatricolporites gamerroi, Tetracolporopollenites megadolium, Todisporites major, T. minor, Tricolpites reticulatus, Triorites orbiculatus, Tuberculatosporites parvus, Tubulifloridites antipodica, Verrustephanoporites simplex, Gomphrenipollis sp., Klukisporites sp., Margocolporites sp., Myrtaceidites sp., Perfotricolpites sp., Reticulatisporites sp., Retistephanocolpites sp., Undulatisporites sp., Verrucingulatisporites sp.Chiquimil
[123]
Acrostichum palaeoaureum, Blechnum serrulatiformis, Cabomba aff. caroliniana, Caesalpinia aff. stuckerti, Cedrela fissiliformis, Ficus tressensii, Lycopodiella aff. cernua, Mayaca aff. fluviatilis, Ranunculodendron anzoteguiae, Sapium haematospermoides, Schinus herbstii, Equisetum sp.Palo
Pintado
[124][125][126]

Correlations
Huayquerian correlations in South America
FormationCerro AzulItuzaingóParanáCamachoRaigónAndalhualaChiquimilLas FloresMaimaráPaloPebasMuyuRosaSaldungaraySalicasUrumacoMap
BasinColoradoParanáHualfínTontalAndesSaltaAmazonHuasiAltiplanoBAVelascoFalcón
Huayquerian (South America)
Country Argentina Uruguay Argentina Brazil
 Peru		 Bolivia Argentina Venezuela
Cardiatherium
Lagostomus
Macroeuphractus
Proeuphractus
Pronothrotherium
Pseudotypotherium
Thylacosmilus
Xotodon
Macraucheniidae
Primates
Rodents
Reptiles
Birds
Terror birds
Flora
EnvironmentsAeolian-fluvialFluvio-deltaicFluvialFluvio-lacustrineFluvialFluvio-lacustrineFluvio-deltaic
Huayquerian volcanoclastics

Huayquerian fauna

Huayquerian flora
VolcanicYes

Notes and references

Notes
  1. Former name of Ituzaingó Formation
  2. Former name of Cerro Azul Formation
  3. In later research considered part of the Camacho Formation[19]
  4. In later research considered part of the Camacho Formation[19]

References
  1. "Huayquerian". Fossilworks. Retrieved 2019-03-21.
  2. Forasiepi et al., 2011, p.136
  3. Garrido et al., 2017, p.54
  4. Muruaga, 2001, p.30
  5. Le Roux et al., 2006, p.46
  6. Verde, 2002, p.21
  7. Pérez, 2012, p.11
  8. Goin et al., 2000, p.101
  9. Verzi et al., 2008, p.146
  10. Morton & Herbst, s.a., p.1
  11. Le Roux et al., 2006, p.49
  12. Cerdeño et al., 2019, p.12
  13. Czaplewski & Campbell, 2017
  14. Frailey, 1986, p.4
  15. García López & Barbot, 2015, p.315
  16. Franco et al., 2013, p.44
  17. Vezzosi, 2015, p.27
  18. Kiyu Formation at Fossilworks.org
  19. Verde, 2002, p.20
  20. Romero et al., 1998, p.60
  21. Pujos et al., 2012, p.2
  22. Marshall & Sempere, 1991, p.636
  23. Córdova Aguilar, 2009, p.26
  24. Morton & Herbst, 2007, p.154
  25. Marshall & Sempere, 1991, p.637
  26. Finger et al., 2007, p.13
  27. Reguero et al., 2015, p.60
  28. Martín Pérez, 2013, p.51
  29. Brea et al., 2013, p.28
  30. Antoine et al., 2016, p.56
  31. Wesselingh et al., 2006, p.304
  32. González & Abascal, 2012, p.129
  33. Brand et al., 2011
  34. Solíz Mundaca, 2018, p.14
  35. Baldellón et al., 1994, p.241
  36. Tófalo & Morrás, 2009, p.683
  37. Marshall & Sempere, 1991
  38. Brandoni et al., 2012, p.10
  39. Cozzuol, 2006, p.190
  40. Linares, 2004, p.16
  41. San Carlos East at Fossilworks.org
  42. Forasiepi et al., 2016, p.11
  43. Cerro Ballena at Fossilworks.org
  44. Bahía Inglesa Unit 2 at Fossilworks.org
  45. Caldera, Bahia Inglesa Unit 1 at Fossilworks.org
  46. Mina Fosforita at Fossilworks.org
  47. Arazati II at Fossilworks.org
  48. Puerto Arazati Beach at Fossilworks.org
  49. Playa Kiyú at Fossilworks.org
  50. Punta Jesus María at Fossilworks.org
  51. Fernicola et al., 2018, p.236
  52. Madozzo Jaén et al., 2018
  53. Sostillo et al., 2014
  54. Laguna Chillhué at Fossilworks.org
  55. Bajo Giuliani at Fossilworks.org
  56. El Guanaco at Fossilworks.org
  57. Telen at Fossilworks.org
  58. Madozzo Jaén & Pérez, 2017
  59. Chiquimil A at Fossilworks.org
  60. Barrancas del Rio Parana at Fossilworks.org
  61. Rio Parana at Fossilworks.org
  62. Salinas Grandes de Hidalgo at Fossilworks.org
  63. De Iullis et al., 2011
  64. Brandoni, 2014
  65. Paraná, Pueblo Brugo to Diamante, Ituzaingó Fm. at Fossilworks.org
  66. Colonia La Celina at Fossilworks.org
  67. Ciudad de Paraná at Fossilworks.org
  68. Playa Sánchez at Fossilworks.org
  69. Prothero et al., 2014
  70. Frailey & Campbell Jr., 2012
  71. LACM loc.6218 at Fossilworks.org
  72. LACM 5159 at Fossilworks.org
  73. VF2 at Fossilworks.org
  74. Bonini et al., 2017
  75. Abello et al., 2015
  76. Maimará at Fossilworks.org
  77. Achiri at Fossilworks.org
  78. Quebrada Pajaritos at Fossilworks.org
  79. El Cajón Valley at Fossilworks.org
  80. Muyu Huasi at Fossilworks.org
  81. Reguero et al., 2015
  82. Cerro La Matanza at Fossilworks.org
  83. Villa Urquiza at Fossilworks.org
  84. Collareta et al., 2017
  85. Marx et al., 2017
  86. Barrancas del Río Chico at Fossilworks.org
  87. Quehua at Fossilworks.org
  88. Arazati km 95.750 at Fossilworks.org
  89. Rosa Pata at Fossilworks.org
  90. Grünbein Cantera Seminario Level 1 at Fossilworks.org
  91. Grünbein Cantera Relleno Sanitario Level 1 at Fossilworks.org
  92. Grünbein Cantera Relleno Sanitario Level 2 at Fossilworks.org
  93. Barrancas de Sarmiento Level 2 at Fossilworks.org
  94. Caleufú at Fossilworks.org
  95. Salicas Formation, Sierra de Velasco, La Rioja at Fossilworks.org
  96. El Degolladito - Site 2 at Fossilworks.org
  97. Kerber et al., 2017
  98. Kerber et al., 2016
  99. Rio Acre at Fossilworks.org
  100. Niteroi at Fossilworks.org
  101. Talismã, upper Purus River at Fossilworks.org
  102. Patos, Rio Acre at Fossilworks.org
  103. Patos at Fossilworks.org
  104. Olivares et al., 2016
  105. Rincón et al., 2018
  106. Rincón et al., 2015
  107. Campo El Mamon at Fossilworks.org
  108. Urumaco, Biozona 3 Loc 73-FU at Fossilworks.org
  109. Urumaco, Biozona 3 Loc 50-FU at Fossilworks.org
  110. Urumaco, Biozona 3 Loc 82-FU at Fossilworks.org
  111. Urumaco, Biozona 3 Loc 26-FU at Fossilworks.org
  112. Valle de Santa María at Fossilworks.org
  113. El Morro at Fossilworks.org
  114. Quebrada Salta, southern area of the Calchaquí Valley at Fossilworks.org
  115. South of Rio Branco at Fossilworks.org
  116. West of El Picacho at Fossilworks.org
  117. 4 km NE of Campo El Mamon at Fossilworks.org
  118. Barranca de Los Loros at Fossilworks.org
  119. Cerro Bautista at Fossilworks.org
  120. Capilla Narbona at Fossilworks.org
  121. Barrancas de San Juan at Fossilworks.org
  122. Arroyo Ensenada valley at Fossilworks.org
  123. Vallecito at Fossilworks.org
  124. Anzótegui & Horn, 2011
  125. Calchaquí at Fossilworks.org
  126. Calchaquí II at Fossilworks.org

Bibliography
Huayquerías Formation
Andalhuala Formation
Camacho Formation
Cerro Azul Formation
Chiquimil Formation
Las Flores Formation, Sierra del Tontal
Içá Formation
Iñapari Formation
India Muerta Formation
Ituzaingó Formation
Madre de Dios Formation
Maimará Formation
Mauri & Muyu Huasi Formations
Miramar Formation
El Morterito Formation
Navidad Formation
Palo Pintado Formation
Paraná Formation
Pebas Formation
Piquete Formation
Pisco Formation
Quehua Formation
Raigón Formation
Saldungaray Formation
Salicas Formation
Solimões Formation
Urumaco Formation
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