Laventan

The Laventan (Spanish: Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the Mayoan age.[1][2]

Etymology

The age is named after the Miocene Lagerstätte La Venta, where a rich biodiversity from the Middle Miocene has been recovered from the Honda Group.[2]

Formations

Locations of Laventan formations

Formation bold is type	Country	Basin	Notes
Honda Group	Colombia	Upper Magdalena Valley	[3]
Honda Group	Bolivia	Quebrada Honda Basin	[4]
Aisol Formation	Argentina	San Rafael Block	[5]
Bahía Inglesa Formation	Chile	Caldera Basin	[6]
Capadare Formation	Venezuela	Falcón Basin	[7]
Choquecota Formation	Bolivia	Altiplano Basin	[8]
Cura-Mallín Group	Chile	Cura-Mallín Basin	[9]
Ipururo Formation	Peru	Ucayali Basin	[10]
Paraná Formation	Argentina	Paraná Basin	[11]
Pebas Formation	Brazil Colombia Ecuador Peru	Amazon Basin	[10][12]
Pisco Formation	Peru	Pisco Basin	[13]
Santa Inés Formation	Venezuela	Eastern Venezuela Basin	[14]
Sincelejo Formation	Colombia	Sinú-San Jacinto Basin	[15]
Socorro Formation	Venezuela	Falcón Basin	[16]
Urumaco Formation	Venezuela	Falcón Basin	[17]

Fossil content

Group	Fossils	Formation	Notes
Mammals	lesser bulldog bat (Noctilio albiventris), LaVal's disk-winged bat (Thyroptera lavali), Spix's disk-winged bat (Thyroptera tricolor), Acarechimys minutissimus, Anachlysictis gracilis, Anadasypus hondanus, Aotus dindensis, Boreostemma acostae, B. gigantea, Brievabradys laventensis, Cebupithecia sarmientoi, Dukecynus magnus, Eodolichotis elachys, E. maddeni, Glossotheriopsis pascuali, Granastrapotherium snorki, Hilarcotherium castanedaii, Hondadelphys fieldsi, Hondathentes cazador, Huilabradys magdaleniensis, Huilatherium pluriplicatum, Kiotomops lopezi, Lagonimico conclucatus, Lycopsis longirostrus, Megadolodus molariformis, Micodon kiotensis, Micoureus laventicus, Microscleromys cribiphilus, M. paradoxalis, Microsteiromys jacobsi, Miocallicebus villaviejai, Miocochilius anomopodus, Mohanamico hershkovitzi, Mormopterus colombiensis, Nanoastegotherium prostatum, Neoglyptatelus originalis, Neonematherium flabellatum, Neoreomys huilensis, Neotamandua borealis, Notonycteris magdalenensis, N. sucharadeus, Nuciruptor rubricae, Pachybiotherium minor, Palynephyllum antimaster, Patasola magdalenae, Pedrolypeutes praecursor, Pericotoxodon platignathus, Pithiculites chenche, Potamops mascahehenes, Potamosiren magdalenensis, Prodolichotis guerreroi, P. pridiana, Prolicaphrium sanalfolsensis, Prothoatherium colombianus, Pseudoprepotherium confusum, Rhodanodolichotis antepridiana, R. vucetichae, Ricardomys longidens, Saimiri annectens, S. fieldsi, Scleromys colombianus, S. schurmanni, Scirrotherium hondaensis, Stirtonia tatacoensis, S. victoriae, Thylamys colombianus, Thylamys minutus, Thyroptera robusta, Villarroelia totoyoi, Xenastrapotherium kraglievichi, Cochilius sp., Diclidurus sp., Eumops sp., Hapalops sp., Olenopsis sp., Theosodon sp., ?Steiromys sp., Dolichotinae sp., ?Echimyidae sp., Megalonychidae sp., Megatheriinae sp., Nothrotheriinae sp., Dasypodidae sp., Leontiniidae sp., Proterotheriidae indet., Toxodontidae sp.	Honda Group Colombia	[18][19][20] [21][22][23] [24][25][26] [27][28][29] [30][31][32] [33][34][35] [36][37][38] [39][40][41] [42][43][44] [45][46][47] [48][49][50] [51][52][53] [54][55][56] [57][58][59] [60][61][62] [63][64][65] [66][67][68] [69][70][71] [72][73][74] [75][76][77] [78][79][80]
	Acyon myctoderos, Australogale leptognathus, Chimeralestes ambiguus, Guiomys unica, Hapalops angustipalatus, Hemihegetotherium trilobus, Hiskatherium saintandrei, Hondalagus altiplanensis, Lakukullus anatisrostratus, Mesoprocta hypsodus, Miocochilius federicoi, Palaeothentes relictus, P. serratus, Propalaehoplophorus andinus, Quebradahondomys potosiensis, Acarechimys sp., cf. Paratrigodon sp., Prolagostomus sp., cf. Prozaedyus sp., Lagostominae indet., Mesotheriinae indet., Borhyaenidae indet., Caenolestidae indet., Capromyidae indet., Caviidae indet., Chinchillidae indet., Echimyidae indet., Euphractinae indet., Macraucheniidae indet., Octodontidae indet., Proterotheriidae indet., ? Xenastrapotherium indet.	Honda Group Bolivia	[81][82] [83][84] [85]
	Nesodon taweretus, cf. Astrapotherium sp., cf. Theosodon sp., Lomaphorini indet., Mylodontinae indet., Propalaehoplophorinae indet., Toxodontidae indet.	Aisol	[86][87]
	Microtypotherium choquecotense, Prozaedyus sp., Chinchillidae indet., Glyptodontidae indet., Hegetotheriidae indet.	Choquecota	[88]
	Heteropsomyinae (aff. Acarechimys) sp., Dasyproctidae (aff. Alloiomys) sp., Prolagostomus sp., Caviomorpha indet., Dasypodidae indet., Typotheria indet.	Cura-Mallín	[89]
	Drytomomys cf. aequatorialis, Granastrapotherium cf. snorki, Miocochilius anomopodus, Neoglyptatelus originalis, Parapropalaehoplophorus septentrionalis, Pericotoxodon cf. platignathus, Potamarchus murinus, cf. Theosodon sp., Acarechimys sp., Boreostemma sp., Megathericulus sp., cf. Tetramerorhinus sp., Urumacotherium sp., Xenastrapotherium sp., Borhyaenoidea indet., Delphinida indet., Glyptodontidae indet., Macraucheniidae indet., Megalonychidae indet., Mylodontidae indet., Octodontoidea indet., Potamarchinae indet., Uruguaytheriinae indet.	Ipururo	[90][91][92] [93][94][95]
	Pontistes rectifrons, Prionodelphis rovereti, Properiptychus argentinus, Protautoga longidens, cf. Balaenoptera sp., Chrysophrys sp., Dioplotherium sp., Balaenidae indet., Physeteridae indet., Sciaenidae indet., ?Sparidae indet.	Paraná	[96]
	Dinomyidae indet., ? Octodontoidea indet.	Pebas	[97]
	Boreostemma venezolensis	Santa Inés	[98]
	Neoglyptatelus sincelejanus	Sincelejo	[15][99]
	Mirandabradys socorrensis, Megatheriidae indet.	Socorro	[100][101]
	cf. Olenopsis sp., Mylodontidae indet.	Urumaco	[102]
Birds	Aramus paludigrus, Galbula hylochoreutes, Hoazinoides magdalenae	Honda Group Colombia	[103][104]
	Psilopteridae indet.	Aisol	[87]
	Pygoscelis calderensis, Spheniscus megaramphus, S. urbinai, Pelagornis sp., aff. Thalassarche sp.	Bahía Inglesa	[105]
	Pelagornis cf. chilensis	Capadare	[106]
Reptiles & amphibians	cane toad (Bufo marinus), Chelus colombiana, Chelonoidis hesterna, Colombophis portai, Dracaena colombiana, ?Eunectes stirtoni, Podocnemis medemi, Gryposuchus colombianus, Langstonia huilensis, Purussaurus neivensis, Balanerodus logimus, Charactosuchus fieldsi, Mourasuchus atopus, Eocaiman sp., Gavialis sp., Sebecus sp., Alligatoridae indet.	Honda Group Colombia	[107][108][109] [110][111][112] [113][114][115] [116][117][118] [119][120][121]
	Crocodylia sp.	Aisol	[87]
	Bairdemys thalassica	Capadare	[122]
	Barinasuchus arveloi	Ipururo	[90]
	Caiman wannlangstoni, Gnatusuchus pebasensis, Gryposuchus pachakamue, Purussaurus neivensis, Kuttanacaiman iquitosensis, Mourasuchus atopus, Paleosuchus sp., Podocnemis sp., Gavialoidea indet.	Pebas	[123] [97][124]
	Colombophis spinosus, Ikanogavialis gameroi, Caiman sp., Mourasuchus sp., Purussaurus sp., Thecachampsa sp., Alethinophidia indet., Boinae indet.	Socorro	[100][125] [126]
	Globidentosuchus cf. brachyrostris, Caiman sp., Mourasuchus sp., Purussaurus sp.	Urumaco	[127]
Fishes & crustaceans	Colossoma macropomum, Lepidosiren paradoxa, cf. Acanthicus, Arapaima sp., Brachyplatystoma cf. B. vaillanti, Brachyplatystoma promagdalena, Sylviocarcinus piriformis, cf. Corydoras sp., Hoplias sp., cf. Hoplosternum sp., Hydrolycus sp., Phractocephalus hemioliopterus, Serrasalmus sp., Pygocentrus sp., or Pristobrycon sp. (cf. Myletes sp.), Ariidae indet., Characidae cf. Tetragonopterinae, Cichlidae indet., Doradidae indet., Loricariidae indet., Potamotrygonidae indet.	Honda Group Colombia	[128][129] [130][131]
	Megalodon, Carcharias acutissima, Carcharodon hastalis, Galeocerdo aduncus, Hemipristis serra, Carcharhinus sp., Heterodontus sp., Squalus sp., Squatina sp., Dasyatidae indet., Myliobatoidea indet., Potamotrygonidae indet.	Paraná	[96]
	cf. Hydrolycus sp., Leporinus sp., Potamotrygon sp., Pristis sp., Anostomidae indet.	Pebas	[97]
	Megalodon, Aetobatus arcuatus, Hemipristis serra, Mugil cephalus, Carcharhinus sp., Citharichthys sp., Galeocerdo sp., Rhinoptera sp., Sphyrna sp., Symphurus sp.	Socorro	[132]
Insects	Macroteleia yaguarum, Sycorax peruensis	Pebas	[133]
Flora	Astroniumxylon parabalansae, Piptadenioxylon paraexcelsa, Solanumxylon paranensis	Paraná	[134]

Correlations

The Laventan (13.8 to 11.8 Ma) correlates with:

NALMA
- latest Barstovian (15.97-13.65 Ma)[135]
- early Clarendonian (13.65-10.3 Ma)[136]
Californian ages
- Luisian (15.5-13.5 Ma)
ELMA - Astaracian (15.97-11.608 Ma)[137]
CPS[138]
- Badenian (13.65-12.7 Ma)[139]
- Sarmatian (12.7-11.608 Ma)[140]
New Zealand stratigraphy - Southland epoch (15.9-10.92 Ma)[141]
- Lillburnian (15.1-12.98 Ma)[142]
- early Waiauan (12.98-10.92 Ma)[143]
Australian ages
- Bairnsdalian (15.0-10.5 Ma)
Japanese ages
- Tozawan (15.97-13.5 Ma)
NMAC - Tunggurian (13.65-11.1 Ma)[144]

Laventan correlations in South America
Formation	Honda	Honda	Aisol	Cura-Mallín	Pisco	Ipururo	Pebas	Capadare	Inés	Paraná	Map
Basin	VSM	Honda	San Rafael	Caldera	Pisco	Ucayali	Amazon	Falcón	Venezuela	Paraná	Laventan (South America)
Country	Colombia	Bolivia	Argentina	Chile	Peru			Venezuela		Argentina
Boreostemma
Hapalops
Miocochilius
Theosodon
Xenastrapotherium
Mylodontidae
Sparassodonta
Primates
Rodents
Birds
Terror birds
Reptiles
megalodon
Flora
Insects
Environments	Fluvial			Fluvio-deltaic		Fluvio-lacustrine		Fluvio-deltaic		Fluvial	Laventan volcanoclastics Laventan fauna Laventan flora
Volcanic	Yes										Laventan volcanoclastics Laventan fauna Laventan flora

References

Paleo Database: Laventan
Madden et al., 1997
Villarroel et al., 1996, p.54
Marshall & Sempere, 1991, p.635
Forasiepi et al., 2011, p.149
Le Roux et al., 2016, p.46
Penín & Villarroel, 2005, p.138
Marshall & Sempere, 1991, p.636
Utgé et al., 2009
Antoine et al., 2016, p.56
Martín Pérez, 2013, p.51
Wesselingh et al., 2006, p.304
Brand et al., 2011
Simpson, 1947, p.2
Villarroel & Clavijo, 2005, p.348
Linares, 2004, p.5
Linares, 2004, p.16
Croft, 2007, p.299
Croft, 2007, p.300
Croft, 2007, p.301
Croft, 2007, p.302
Croft, 2007, p.303
Meldrum & Kay, 1997
Organ & Lemelin, 2011
Rosenberger et al., 1991
Setoguchi & Rosenberger, 1988
Setoguchi, 1985
Takai et al., 2001
Tejedor, 2013, p.22
Defler, 2004, p.32
Cebupithecia sarmientoi at Fossilworks.org
Miocallicebus villaviejai at Fossilworks.org
Neosaimiri annectens at Fossilworks.org
Stirtonia tatacoensis at Fossilworks.org
Stirtonia victoriae at Fossilworks.org
Vallejo Pareja et al., 2015
Pardo Jaramillo, 2010
Granastrapotherium snorki, Sebecus sp. at Fossilworks.org
Villarroel, 2000, p.118
Kay & Madden, 1997, p.171
Neonematherium flabellatum at Fossilworks.org
Brievabradys laventensis at Fossilworks.org
Huilabradys magdaleniensis at Fossilworks.org
Czaplewski et al., 2003, p.278
Neotamandua borealis at Fossilworks.org
Pseudoprepotherium confusum at Fossilworks.org
Anadasypus hondanus at Fossilworks.org
Boreostemma acostae at Fossilworks.org
Boreostemma gigantea at Fossilworks.org
Neoglyptatelus originalis at Fossilworks.org
Pedrolypeutes praecursor at Fossilworks.org
Scirrotherium hondaensis at Fossilworks.org
Prolicaphrium sanalfolsensis at Fossilworks.org
Prothoatherium colombianus at Fossilworks.org
Duke Locality 33 at Fossilworks.org
Villarroelia totoyoi at Fossilworks.org
Villarroel, 1997
Villarroel et al., 1996, p.63
Hoffstetter, 1971, p.40
Suárez et al., 2015
Marshall, 1976
Hondadelphys fieldsi at Fossilworks.org
Pachybiotherium minor at Fossilworks.org
Thylamys colombianus at Fossilworks.org
Thylamys minutus at Fossilworks.org
Hondathentes cazador at Fossilworks.org
Pithiculites chenche at Fossilworks.org
Neoreomys huilensis at Fossilworks.org
Rhodanodolichotis antepridiana at Fossilworks.org
Scleromys colombianus at Fossilworks.org
Scleromys schurmanni at Fossilworks.org
Prodolichotis guerreroi at Fossilworks.org
UCMP V4936 at Fossilworks.org
Eodolichotis elachys at Fossilworks.org
Eodolichotis maddeni at Fossilworks.org
Rhodanodolichotis vucetichae at Fossilworks.org
Takai et al., 1991
Kiotomops lopezi at Fossilworks.org
Potamops mascahehenes at Fossilworks.org
Thyroptera robusta at Fossilworks.org
Engelman et al., 2016
Pujos et al., 2014
Hondalagus at Fossilworks.org
Quebrada Honda at Fossilworks.org
Engelman et al., 2018
Forasiepi et al., 2015
Cerro Aisol at Fossilworks.org
Choquecota at Fossilworks.org
Cerro los Pinos at Fossilworks.org
IN-DTC-20 at Fossilworks.org
IN-DTC-28 at Fossilworks.org
IN-DTC-32 at Fossilworks.org
Río Sepa at Fossilworks.org
SEP 005 at Fossilworks.org
IN 008 at Fossilworks.org
Pueblo Brugo to Diamante at Fossilworks.org
CTA-45 at Fossilworks.org
Banks of the Rio Guere at Fossilworks.org
Calle Fria-Segovia at Fossilworks.org
South East of 'Cerro los Chivatos' at Fossilworks.org
Mirandabradys Urumaco 01 at Fossilworks.org
Biozona 1 Loc 44-FU at Fossilworks.org
Rasmussen, 1997
Miller, 1953
Los Dedos at Fossilworks.org
Zumbador Cave at Fossilworks.org
Estes & Wassersug, 1963
Gryposuchus colombianus at Fossilworks.org
Sebecus huiensis at Fossilworks.org
Dracaena colombiana at Fossilworks.org
Purussaurus neivensis at Fossilworks.org
Charactosuchus fieldsi at Fossilworks.org
Mourasuchus atopus at Fossilworks.org
UCMP Locality V4524 at Fossilworks.org
Balanerodus logimus at Fossilworks.org
Eocaiman at Fossilworks.org
Cadena et al., 2008, p.1206
De la Fuente et al., 2013, p.102
Geochelone hesterna at Fossilworks.org
Hsiou et al., 2010
(in Spanish) Fauna de La Venta - serpientes
Ferreira et al., 2015
Salas Gismondi et al., 2015
IQ114 at Fossilworks.org
Quebrada Honda at Fossilworks.org
East of Capirote, road to Quebrada Honda at Fossilworks.org
Sur Quebrada Bejucal at Fossilworks.org
(in Spanish) Fauna de La Venta - flora, cangrejos y peces
Kay & Madden, 1997, p.185
Lundberg & Chernoff, 1992, p.5
Brachyplatystoma promagdalena at Fossilworks.org
Socorro at Fossilworks.org
Amazonian amber at Fossilworks.org
Toma Vieja at Fossilworks.org
Barstovian at Fossilworks.org
Clarendonian at Fossilworks.org
Astaracian at Fossilworks.org
Central Paratethys Stages at Fossilworks.org
Badenian at Fossilworks.org
http://fossilworks.org/bridge.pl?action=displayInterval&interval_no=1045 Sarmatian] at Fossilworks.org
Southland epoch at Fossilworks.org
Lillburnian at Fossilworks.org
Waiauan at Fossilworks.org
Tunggurian at Fossilworks.org

Bibliography

Laventan

Madden, R.H.; J. Guerrero; R.F. Kay; J.J. Flynn; C.C. Swisher III, and A.H. Walton. 1997. The Laventan Stage and Age, 499–519. Kay, R.F., Madden, R.H., Cifelli, R.L., Flynn, J.J. (eds.), Vertebrate Paleontology in the Neotropics. The Miocene Fauna of La Venta, Colombia: Washington D.C., Smithsonian Institution Press. Accessed 2019-02-28.

Honda Group, Colombia

Cadena, Edwin; Carlos Jaramillo, and María E. Páramo. 2008. New material of Chelus colombiana (Testudines; Pleurodira) from the Lower Miocene of Colombia. Journal of Vertebrate Paleontology 28. 1206–1212. Accessed 2017-05-04.
Croft, Darin A. 2007. The Middle Miocene (Laventan) Quebrada Honda Fauna, southern Bolivia and a description of its Notoungulates. Palaeontology 50. 277–303.
Czaplewski, Nicolas J.; Masanaru Takai; Tiffany M. Naeher; Nobuo Shigehara, and Takeshi Setoguchi. 2003. Additional bats from the middle Miocene La Venta fauna of Colombia. Revista Académica Colombiana de Ciencias Naturales XXVII. 263–282. Accessed 2017-05-04.
Defler, Thomas. 2004. Historia natural de los primates colombianos, 1–613. Universidad Nacional de Colombia. Accessed 2017-05-04.
Estes, Richard, and Richard Wassersug. 1963. A Miocene toad from Colombia, South America. Breviora 193. 1–13. Accessed 2017-05-04.
De la Fuente, Marcelo; Juliana Sterli, and Ignacio Maniel. 2013. Origin, Evolution and Biogeographic History of South American Turtles, 1–168. Springer Science & Business Media. Accessed 2017-05-04.
Hoffstetter, Robert. 1971. Los vertebrados cenozóicos de Colombia: yacimientos, faunas, problemas planteados, 37–62. Universidad Nacional de Colombia. Accessed 2017-05-04.
Hsiou, Annie S.; Adriana M. Albino, and Jorge Ferigolo. 2010. Reappraisal of the south American Miocene snakes of the genus Colombophis, with description of a new species. Acta Palaeontologica Polonica 55. 365–379.
Kay, Richard F., and Richard H. Madden. 1997. Mammals and rainfall: paleoecology of the middle Miocene at La Venta (Colombia, South America). Journal of Human Evolution 32. 161–199. Accessed 2017-05-04.
Lundberg, John G., and Barry Chernoff. 1992. A Miocene Fossil of the Amazonian Fish Arapaima (Teleostei, Arampaimidae) from the Magdalena River Region of Colombia - Biogeographic and Evolutionary Implications. Biotropica 24. 2–16. Accessed 2017-05-04.
Marshall, Larry G. 1976. New Didelphine Marsupials from the La Venta Fauna (Miocene) of Colombia, South America. Journal of Paleontology 50. 402–418.
Meldrum, D.J., and R.F. Kay. 1997. <407::AID-AJPA8>3.0.CO;2-R Nuciruptor rubricae, a new pitheciin seed predator from the Miocene of Colombia. American Journal of Physical Anthropology 102. 407–427. Accessed 2017-05-04.
Miller, Alden H. 1953. A fossil Hoatzin from the Miocene of Colombia. Auk 70. 484–495. Accessed 2017-05-04.
Organ, Jason M., and Pierre Lemelin. 2011. Tail Architecture and Function of Cebupithecia sarmientoi, a Middle Miocene Platyrrhine from La Venta, Colombia. The Anatomical Record 294. 2013–2023.
Pardo Jaramillo, Mauricio. 2010. Reporte de un nuevo ejemplar de Granastrapotherium snorki en el Valle Superior del Magdalena, Desierto de la Tatacoa, Huila. Colombia. Revista Académica Colombiana de Ciencias Naturales 34. 253–256. Accessed 2017-05-04.
Rasmussen, Tab. 1997. Vertebrate paleontology in the neotropics - the Miocene fauna of La Venta, Colombia, _. Smithsonian Institution Press.
Rosenberger, A.L.; T. Setoguchi, and W.C. Hartwig. 1991. Laventiana annectens, new genus and species: fossil evidence for the origins of callitrichine New World monkeys. Proceedings of the National Academy of Sciences of the United States of America 88. 2137–2140.
Setoguchi, Takeshi, and Alfred L. Rosenberger. 1988. A fossil owl monkey from La Venta, Colombia. Kyoto University Overseas Research Reports of New World Monkeys 6. 1–6. Accessed 2017-05-04.
Setoguchi, T. 1985. Kondous laventicus, a new ceboid primate from the Miocene of the La Venta, Colombia, South America. Folia Primatol 44. 96–101.
Suárez, Catalina; Analía M. Forasiepi; Franciso J. Goin, and Carlos Jaramillo. 2015. Insights into the Neotropics prior to the Great American Biotic Interchange: new evidence of mammalian predators from the Miocene of Northern Colombia. Journal of Vertebrate Paleontology e1029581. 1–10. Accessed 2017-05-04.
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Takai, Masanaru; Takeshi Setoguchi; Carlos Villarroel A.; Alberto Cadena G., and Nobuo Shigehara. 1991. A New Miocene Molossid Bat from La Venta, Colombia, South America. Memoirs of the Faculty of Science, Kyoto University. Series of Geology and Mineralogy 56. 1–9. Accessed 2017-05-04.
Tejedor, Marcelo F. 2013. Sistemática, evolución y paleobiogeografía de los primates Platyrrhini. Revista del Museo de la Plata 20. 20–39. Accessed 2017-05-04.
Vallejo Pareja, M.C.; J.D. Carrillo; J.W. Moreno Bernal; M. Pardo Jaramillo; D.F. Rodríguez González, and J. Muñoz Duran. 2015. Hilarcotherium castanedaii, gen. et sp. nov., a new Miocene astrapothere (Mammalia, Astrapotheriidae) from the Upper Magdalena Valley, Colombia. Zurich Open Repository and Archive, University of Zurich _. 1–14. Accessed 2017-05-04.
Villarroel A., Carlos S. 2000. Un nuevo Mylodontinae (Xenarthra, Tardigrada) en la Fauna de La Venta, Mioceno de Colombia: el estado actual de la Familia Orophodontidae. Revista de la Academia Colombiana de Ciencias Exactas Físicas y Naturales 24. 117–127. Accessed 2017-05-04.
Villarroel A., Carlos S. 1997. La Estructura de la Dentición Caduca de Huilatherium pluriplicatum, Leontiniidae (Notoungulata) del Mioceno de Colombia. Geología Colombiana 22. 139–149. Accessed 2017-05-04.
Villarroel A., Carlos; Takeshi Setoguchi; Jorge Brieva, and Carlos Macia. 1996. Geology of the La Tatacoa "Desert" (Huila, Colombia): Precisions on the Stratigraphy of the Honda Group, the Evolution of the "Pata High" and the Presence of the La Venta Fauna. Memoirs of the Faculty of Science, Kyoto University. Series of Geology and Mineralogy 58. 41–66. Accessed 2017-05-04.
Wheeler, Brandon. 2010. Community ecology of the Middle Miocene primates of La Venta, Colombia: the relationship between ecological diversity, divergence time, and phylogenetic richness. Primates 51. 131–138.

Honda Group, Bolivia

Croft, Darin A. 2007. The Middle Miocene (Laventan) Quebrada Honda Fauna, southern Bolivia and a description of its Notoungulates. Palaeontology 50. 277–303.
Engelman, Russell K.; Federico Anaya, and Darin A. Croft. 2018. Australogale leptognathus, gen. et sp. nov., a second species of small sparassodont (Mammalia: Metatheria) from the middle Miocene locality of Quebrada Honda, Bolivia. Journal of Mammalian Evolution in press. _. Accessed 2018-10-01.
Engelman, Russell K.; Federico Anaya, and Darin A. Croft. 2016. New palaeothentid marsupials (Paucituberculata) from the middle Miocene of Quebrada Honda, Bolivia, and their implications for the palaeoecology, decline and extinction of the Palaeothentoidea. Journal of Systematic Palaeontology 15. 787–820. Accessed 2019-02-12.
Pujos, François; Gerardo De Iuliis; Bernardino Mamani Quispe, and Ruben Andrade Flores. 2014. Lakukullus anatisrostratus, gen. et sp. nov., a new massive nothrotheriid sloth (Xenarthra, Pilosa) from the middle Miocene of Bolivia. Journal of Vertebrate Paleontology 34. 1243–1248. Accessed 2019-02-12.

Aisol Formation

Forasiepi, Analía M.; Esperanza Cerdeño; Mariano Bond; Gabriela I. Schmidt; Maximiliano Naipauer; Fiona R. Straehl; Agustín G. Martinelli; Alberto C. Garrido, and Mark D. Schmitz and James L. Crowley. 2015. New toxodontid (Notoungulata) from the Early Miocene of Mendoza, Argentina. Paläontologische Zeitschrift 89. 611–634. Accessed 2019-02-13.
Forasiepi, Analía M.; Agustín G. Martinelli; Marcelo S. De la Fuente; Sergio Dieguez, and Mariano Bond. 2011. Paleontology and stratigraphy of the Aisol Formation (Neogene), San Rafael, Mendoza, 135–154. Cenozoic Geology of the Central Andes of Argentina, SCS Publisher, Salta. Accessed 2017-10-01.

Bahía Inglesa Formation

Achurra, L.E.; J.P. Lacassie; J.P. Le Roux; C. Marquadt; M. Belmar; J. Ruiz Del Solar, and S.E. Ishman. 2009. Manganese nodules in the Miocene Bahía Inglesa Formation, north-central Chile: Petrography, geochemistry, genesis and palaeoceanographic significance. Sedimentary Geology 217. 128–139. Accessed 2017-08-15.
Achurra, L.; C. Marquadt; J.P. Lacassie, and J.P. Le Roux. 2003. Nódulos de Mn de la Formación Bahía Inglesa: geoquímica, génesis e implicancias paleoambientales, 1–10. X Congreso Geológico Chileno. Accessed 2017-08-15.
Acosta Hospitaleche, Carolina, and Jhoann Canto. 2005. Primer registro de cráneos asignados a Palaeospheniscus (Aves, Spheniscidae) procedentes de la Formación Bahía Inglesa (Mioceno Medio-Tardío), Chile. Revista Chilena de Historia Natural 78. 489–495. Accessed 2017-08-15.
Benson, Richard D. 2015. A new species of penguin from the late Miocene of Chile, with comments on the stratigraphic range of Palaeospheniscus. Scientific Publications of the Science Museum of Minnesota. New Series 8. 5–22. Accessed 2019-02-13.
Chávez Hoffmeister, Martín Felipe. 2008. La ornitofauna de la Formación Bahía Inglesa, Caldera, Chile (MSc. thesis), 1–165. Universidad Austral de Chile. Accessed 2017-08-15.
Chávez, Martín F. 2007. Sobre la presencia de Paraptenodytes y Palaeospheniscus (Aves: Sphenisciformes) en la Formación Bahía Inglesa, Chile. Revista Chilena de Historia Natural 80. 255–259. Accessed 2017-08-15.
Chávez, Martín. 2007. Spheniscus megaramphus Stucchi et al. 2003 (Aves: Sphenisciformes) en la Formación Bahía Inglesa, Chile. Boletín de la Sociedad Geológica del Perú 102. 101–107. Accessed 2017-08-15.
Le Roux, Jacobus P.; Luciano Achurra; Álvaro Henríquez; Catalina Carreño; Huber Rivera; Mario E. Suárez; Scott E. Ishman; Nicholas D. Pyenson, and Carolina S. Gutstein. 2016. Oroclinal bending of the Juan Fernández Ridge suggested by geohistory analysis of the Bahía Inglesa Formation, north-central Chile. Sedimentary Geology 333. 32–49. Accessed 2017-08-15.
Le Roux, Jacobus P.; Luciano Achurra; Álvaro Henríquez; Catalina Carreño, and Huber Rivera. 2015. Lithostratigraphy, depositional environments and tectonic setting of the Bahía Inglesa Formation west of Copiapó, north-central Chile, 821–824. XIV Congreso Geológico Chileno. Accessed 2017-08-15.
Oyanadel Urbina, Pablo; Jorge Carrillo Briceño; Jaime Villafaña; Victor Castelleto; Cristian Varas; Alex Alballay, and Marcelo Rivadeneira. 2015. Nuevo registro de familias de peces óseos en Formación Bahía Inglesa, 1–4. XIV Congreso Geológico Chileno. Accessed 2017-08-15.
Staig, Felipe; Sebastián Hernández; Patricio López; Jaime A. Villafaña; Cristian Varas; Luis Patricio Soto, and Jorge D. Carrillo Briceño. 2015. Late Neogene Elasmobranch fauna from the Coquimbo Formation, Chile. Revista Brasileira de Paleontologia 18. 261–272. Accessed 2017-08-15.
Suárez, Mario E. 2011. Línea de base paleontológica proyecto Puerto Santa Fe, 1–29. Comuna de Caldera Región de Atacama. Accessed 2017-08-15.
Valenzuela Toro, Ana, and Carolina S. Gutstein. s.a. Mamíferos marinos fósiles (excepto Cetacea) de Formación Bahía Inglesa: Registro, implicancias paleoambientales y perspectivas de estudio. . .. 770–772. Accessed 2017-08-15.

Capadare Formation

Ferreira, Gabriel S.; Ascanio D. Rincón; Andrés Solórzano, and Max C. Langer. 2015. The last marine pelomedusoids (Testudines: Pleurodira): a new species of Bairdemys and the paleoecology of Stereogenyina. PeerJ 3. e1063. Accessed 2019-02-13.
Penín Cendón, José Antonio, and Vanessa Ilia Villarroel Estaba. 2005. Integracrón y actualización geológica del área de Jacura y Capadare, Estado Falcón, 1–199. Universidad Central de Venezuela. Accessed 2017-08-15.

Choquecota and Mauri Formations

Marshall, Larry G., and Thierry Sempere. 1991. The Eocene to Pleistocene vertebrates of Bolivia and their stratigraphic context: A review. Revista técnica de YPFB 12. 631–652. Accessed 2017-08-15.

Cura-Mallín Group

Flynn, John J.; Reynaldo Charrier; Darin A. Croft; Phillip B. Gans; Trystan M. Herriott; Jill A. Wertheim, and André R. Wyss. 2008. Chronologic implications of new Miocene mammals from the Cura-Mallín and Trapa Trapa formations, Laguna del Laja area, south central Chile. Journal of South American Earth Sciences 26. 412–423. Accessed 2017-10-20.
Shockey, Bruce J.; John J. Flynn; Darin A. Croft; Phillip Gans, and André R. Wyss. 2012. New leontiniid Notoungulata (Mammalia) from Chile and Argentina : comparative anatomy, character analysis, and phylogenetic hypotheses. American Museum Novitates 3737. 1–64. Accessed 2019-02-15.
Utgé, Silvana; Andrés Folguera; Vanesa Litvak, and Víctor A. Ramos. 2009. Geología del sector norte de la Cuenca de Cura Mallín en las Lagunas de Epulaufquen, Neuquén. Revista de la Asociación Geológica Argentina 64. 231–248. Accessed 2017-10-01.

Gran Bajo del Gualicho Formation

Reichler, Valeria A. 2010. Estratigrafía y paleontología del Cenozoico marino del Gran Bajo y Salinas del Gualicho, Argentina, y descripción de 17 especies nuevas. Andean Geology 31. 177–219. Accessed 2018-09-11.

Ipururo Formation

Antoine, Pierre-Olivier; M. Alejandra Abello; Sylvain Adnet; Ali J. Altamirano Sierra; Patrice Baby; Guillaume Billet; Myriam Boivin; Ysabel Calderón, and Adriana Candela, Jules Chabain, Fernando Corfu, Darin A. Croft, Morgan Ganerød, Carlos Jaramillo, Sebastian Klaus, Laurent Marivaux, Rosa E. Navarrete, Maëva J. Orliac, Francisco Parra, María Encarnación Pérez, François Pujos, Jean-Claude Rage, Anthony Ravel, Céline Robinet, Martin Roddaz, Julia Victoria Tejada Lara, Jorge Vélez Juarbe, Frank P. Wesselingh, Rodolfo Salas Gismondi. 2016. A 60-million-year Cenozoic history of western Amazonian ecosystems in Contamana, eastern Peru. Gondwana Research 31. 30–59. Accessed 2017-08-15.

Paraná Formation

Cione, Alberto Luis; Daniel Alfredo Cabrera, and María Julia Barla. 2012. Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic. Geobios 45. 167–172. Accessed 2017-10-22.
Martín Pérez, Leandro. 2013. Sistemática, tafonomía y paleoecología de los invertebrados de la Formación Paraná (Mioceno), Provincia de Entre Ríos, Argentina (PhD thesis), 1–398. Universidad Nacional de La Plata. Accessed 2017-08-15.

Pebas Formation

Antoine, Pierre-Olivier; M. Alejandra Abello; Sylvain Adnet; Ali J. Altamirano Sierra; Patrice Baby; Guillaume Billet; Myriam Boivin; Ysabel Calderón, and Adriana Candela, Jules Chabain, Fernando Corfu, Darin A. Croft, Morgan Ganerød, Carlos Jaramillo, Sebastian Klaus, Laurent Marivaux, Rosa E. Navarrete, Maëva J. Orliac, Francisco Parra, María Encarnación Pérez, François Pujos, Jean-Claude Rage, Anthony Ravel, Céline Robinet, Martin Roddaz, Julia Victoria Tejada Lara, Jorge Vélez Juarbe, Frank P. Wesselingh, Rodolfo Salas Gismondi. 2016. A 60-million-year Cenozoic history of western Amazonian ecosystems in Contamana, eastern Peru. Gondwana Research 31. 30–59. Accessed 2017-08-15.
Wesselingh, F.P.; M.C. Hoorn; J. Guerrero; M.E. Räsänen; L. Romero Pittmann, and J. Salo. 2006. The stratigraphy and regional structure of Miocene deposits in western Amazonia (Peru, Colombia and Brazil), with implications for late Neogene landscape evolution. Scripta Geologica 133. 291–322. Accessed 2017-08-15.

Pisco Formation

Altamirano Sierra, Alí J. 2013. Primer registro de pelicano (Aves: Pelecanidae) para el Mioceno tardio de la formacion Pisco, Peru. Bulletin de l'Institut français d'études andines 42. 1–12. Accessed 2017-09-04.
Báez Gómez, Diego A. 2006. Estudio paleoambiental de la formación Pisco:: Localidad Ocucaje. Revista del Instituto de Investigaciones FIGMMG 9. 64–69. Accessed 2017-09-04.
Bianucci, Giovanni; Claudio Di Celma; Mario Urbina, and Olivier Lambert. 2016. New beaked whales from the late Miocene of Peru and evidence for convergent evolution in stem and crown Ziphiidae (Cetacea, Odontoceti). PeerJ 4. e2479. Accessed 2019-02-12.
Brand, Leonard; Mario Urbina; Arthur Chadwick; Thomas J. DeVries, and Raul Esperante. 2011. A high resolution stratigraphic framework for the remarkable fossil cetacean assemblage of the Miocene/Pliocene Pisco Formation, Peru. Journal of South American Earth Sciences 31. 414–425.
Collareta, Alberto; Olivier Lambert; Christian De Muizon; Mario Urbina, and Giovanni Bianucci. 2017. Koristocetus pescei gen. et sp. nov., a diminutive sperm whale (Cetacea: Odontoceti: Kogiidae) from the late Miocene of Peru. Fossil Record 20. 259–278. Accessed 2019-02-12.
Lambert, Olivier; Giovanni Bianucci; Mario Urbina, and Jonathan H. Geisler. 2017. A new inioid (Cetacea, Odontoceti, Delphinida) from the Miocene of Peru and the origin of modern dolphin and porpoise families. Zoological Journal of the Linnean Society 179. 919–946.
Marx, Felix G.; Olivier Lambert, and Christian De Muizon. 2017. A new Miocene baleen whale from Peru deciphers the dawn of cetotheriids. Royal Society Open Science 4. 170560. Accessed 2019-02-12.
Marx, Felix G., and Naoki Kohno. 2016. A new Miocene baleen whale from the Peruvian desert. Royal Society Open Science 3. 160542. Accessed 2019-02-12.
Poma Porras, Orlando; Edgard Horna Santillán, and Raúl Esperante. 2009. Baleen Fósil (Cetacea: mysticeti) en Sedimentos de la Cuenca Marina del Neógeno en la Formación Pisco, al Sur del Perú. Revista de Investigación Universitaria 1. 84–97. Accessed 2017-09-04.
Ramassamy, Benjamin; Olivier Lambert; Alberto Collareta; Mario Urbina, and Giovanni Bianucci. 2018. Description of the skeleton of the fossil beaked whale Messapicetus gregarius: searching potential proxies for deep-diving abilities. Fossil Record 21. 11–32. Accessed 2019-02-11.
Solís Mundaca, Flavio Alejandro. 2018. Bioestratigrafía e implicancias paleoceanográficas de las diatomeas de la sección Cerro Caucato, Formación Pisco, Ica, Peru (MSc. thesis), 1–158. Universidad Peruana Cayetano Heredia. Accessed 2018-09-10.
Stucchi, Marcelo; Steven D. Emslie; Rafael M. Varas Malca, and Mario Urbina Schmitt. 2015a. A new late Miocene condor (Aves, Cathartidae) from Peru and the origin of South American condors. Journal of Vertebrate Paleontology 35. e972507. Accessed 2019-02-13.
Stucchi, Marcelo; Rafael M. Varas Malca, and Mario Urbina Schmitt. 2015b. New Miocene sulid birds from Peru and considerations on their Neogene fossil record in the Eastern Pacific Ocean. Acta Palaeontologica Polonica 61. 417–427. Accessed 2019-02-13.
Stucchi, M. 2007. Los pingüinos de la Formación Pisco (Neógeno), Perú. 4th European Meeting on the Palaeontology and Stratigraphy of Latin America, Cuadernos del Museo Geominero 8. 367–373. Accessed 2017-09-04.

Santa Inés Formation

Simpson, George Gaylord. 1947. A Miocene glyptodont from Venezuela. American Museum Novitates 1368. 1–10.

Sincelejo Formation

Villarroel A., Carlos, and Javier Clavijo. 2005. Los mamíferos fósiles y las edades de las sedimentitas continentales del Neógeno de la Costa Caribe Colombiana. Revista de la Academia Colombiana de Ciencias Exactas Físicas y Naturales 29. 345–356. Accessed 2019-02-04.

Socorro and Urumaco Formations

Carrillo Briceño, Jorge D.; Erin Maxwell; Órangel A. Aguilera; Rodolfo Sánchez, and Marcelo R. Sánchez Villagra. 2015. Sawfishes and Other Elasmobranch Assemblages from the Mio-Pliocene of the South Caribbean (Urumaco Sequence, Northwestern Venezuela). PLoS ONE 10. e0139230.
Linares, Omar J. 2004. Bioestratigrafía de la fauna de mamíferos de las formaciones Socorro, Urumaco y Codore (Mioceno Medio-Plioceno Temprano) de la región de Urumaco, Falcón, Venezuela. Paleobiología Neotropical 1. 1–26. Accessed 2017-08-15.
Quijano Ballesteros, Jhon Richard. 2005. Estudio magnetoestratigráfico en la sección de El Mamón (miembro medio de la Formación Urumaco, Estado Falcón) (MSc. thesis), 1–92. Universidad Simón Bolívar. Accessed 2017-10-27.
Rincón, Ascanio D.; Andrés Solórzano; H. Gregory McDonald, and Marisol Montellano Ballesteros. 2018. Two new megalonychid sloths (Mammalia: Xenarthra) from the Urumaco Formation (late Miocene), and their phylogenetic affinities. Journal of Systematic Palaeontology 17. 409–421.
Rincón, Ascanio D.; H. Gregory McDonald; Andrés Solórzano; Mónica Núñez Flores, and Damián Ruiz Ramoni. 2015. A new enigmatic Late Miocene mylodontoid sloth from northern South America. Royal Society Open Science 2. 140256. Accessed 2019-02-13.