Patriomanis

Patriomanis; Temporal range: 37.2–33.9 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ Late Eocene
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Pholidota
Suborder:	Eupholidota
Superfamily:	Manoidea
Family:	†Patriomanidae
Genus:	†Patriomanis; Emry, 1970
Type species
	Patriomanis americana; Emry, 1970[1]
Synonyms
	P. americana: Patriomanis americanus (Emry, 1970) ;

Patriomanis ("father of pangolins") is an extinct genus of pangolin from extinct family Patriomanidae. It lived during the Eocene of North America and it currently represents the only pangolin known from the Western Hemisphere. The genus contains one species, P. americana, which is known from six specimens, mostly from the Chadronian White River Formation of Montana. It had long digits and a prehensile tail, suggesting that it was arboreal, and its jaw was capable of opening wider than modern pangolins. Its ears and the hair between its scales were also longer than modern pangolins.[2]

Phylogeny

Phylogenetic position of genus Patriomanis within family Patriomanidae.[3][4]

Pholidotamorpha

†Palaeanodonta

Pholidota

†Euromanis

?

†pholidota sp. (BC 16’08)

†Eurotamanduidae

Eupholidota

†Eomanoidea

Manoidea

	Manidae

?	†Necromanis

†Patriomanidae

	†Cryptomanis

†Patriomanis	†Patriomanis americana

sensu stricto

(Pholidota sensu lato)

References

R. J. Emry. (1970.) "A North American Oligocene pangolin and other additions to the Pholidota." Bulletin of the American Museum of Natural History 142(6)
Gaudin, T. J.; Emry, R. J.; Morris, J. (2016). "Skeletal anatomy of the North American pangolin Patriomanis americana (Mammalia, Pholidota) from the Latest Eocene of Wyoming (USA)". Smithsonian Contributions to Paleobiology (98): vii-102. doi:10.5479/si.1943-6688.98.
Gaudin, Timothy (2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis" (PDF). Journal of Mammalian Evolution. Heidelberg, Germany: Springer Science+Business Media. 16 (4): 235–305. doi:10.1007/s10914-009-9119-9. S2CID 1773698.
Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis". Journal of Vertebrate Paleontology. 32 (5): 983–1001. doi:10.1080/02724634.2012.694319. ISSN 0272-4634. S2CID 86059673.

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[1] R. J. Emry. (1970.) "A North American Oligocene pangolin and other additions to the Pholidota." Bulletin of the American Museum of Natural History 142(6)

[2] Gaudin, T. J.; Emry, R. J.; Morris, J. (2016). "Skeletal anatomy of the North American pangolin Patriomanis americana (Mammalia, Pholidota) from the Latest Eocene of Wyoming (USA)". Smithsonian Contributions to Paleobiology (98): vii-102. doi:10.5479/si.1943-6688.98.

[web2.utc.edu-3] Gaudin, Timothy (2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis" (PDF). Journal of Mammalian Evolution. Heidelberg, Germany: Springer Science+Business Media. 16 (4): 235–305. doi:10.1007/s10914-009-9119-9. S2CID 1773698.

[KondrashovAgadjanian2012-4] Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis". Journal of Vertebrate Paleontology. 32 (5): 983–1001. doi:10.1080/02724634.2012.694319. ISSN 0272-4634. S2CID 86059673.