Texacephale

Texacephale langstoni dome discovered in the Aguja Formation

The holotype specimen of Texacephale, LSUMNS 20010, is composed of fused frontals and parietals. A second specimen, LSUMNS 20012, is composed of an incomplete frontoparietal dome. It was found in the same WPA quarries that produced Agujaceratops, and may have been exavated and tossed aside, being mistaken for a rock or concretion, before being picked up decades later.[1] According to the team, the fossilized dome of the animal possessed five to six vertical flanges on each lateral side, connecting it with the postorbital bone. The team interpreted these structures as interlocking "gears" that would help deal with stress on the bone during head-butting,[2] a hypothetical behavior that had earlier been challenged by other authors.[3][4]

Texacephale was, like other pachycephalosaurids, a small, bipedal herbivore. The thickened dome, formed by the fused frontals and parietals, with contributions from other bones of the skull, would have meant that the brain was covered by several inches of solid bone. The dome is likely to have been used in head-butting contests over mates and/or territory, similar to modern head-butting species such as bighorn sheep, cape buffalo, and musk oxen. Damage to the skull bone may have been sustained during combat.

In contrast to today's dry, desert environment, the Big Bend paleoenvironment was a wet, coastal lowland on the edge of the Western Interior Seaway during the Campanian. Texacephale shared its environment with Agujaceratops mariscalensis, a ceratopsid. Tyrannosaurids and the giant crocodilian Deinosuchus rugosus were also present in the Aguja environment, and were likely to have been predators.

Cladogram after Longrich, Sankey and Tanke (2010).[1]

• Timeline of pachycephalosaur research