Diplocynodon

Diplocynodon is an extinct genus of alligatoroid that lived during the Paleocene to middle Miocene in Europe. It looked very similar to the modern caiman in that it was small and had bony armour scutes covering its neck, back, belly, and tail. The longest Diplocynodon recovered was 4 feet in length and probably fed on small fish, frogs, and took insects when young.

Diplocynodon
Temporal range: Paleocene–Middle Miocene
Diplocynodon ratelii
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Crocodilia
Subfamily: Diplocynodontinae
Genus: Diplocynodon
Pomel, 1847
Species
  • D. dalpiazi Del Favero, 1999
  • D. darwini (Ludvig, 1877)
  • D. elavericus Martin, 2010
  • D. gervaisi
  • D. hantoniensis (Wood, 1846)
  • D. muelleri (Kälin, 1936)
  • D. ratelii Pomel, 1847 (type)
  • D. tormis
  • D. ungeri (Prangner, 1845)
  • ?D. deponiae (Frey, Laemmert & Riess, 1987)[1]
Synonyms

In the nineteenth century, D. steineri was named from Styria, Austria and D. styriacus was named from Austria and France. A third Austrian species, Enneodon ungeri, was placed in its own genus. The Austrian and French species of Diplocynodon were synonymized with E. ungeri in 2011, and because the name Diplocynodon has priority over Enneodon, the species is now called D. ungeri. [2]

Well preserved specimens have been found in the Messel Pit and the Geiseltal lignite deposit in Germany. Most articulated Diplocynodon specimens from these localities contain gastroliths. In the Eocene epoch, the German sites were either a swampy freshwater lake (Messel Pit) or a peat bog swamp (Geiseltal).

Species

Species
Species Age Location Unit Notes Images

D. dalpiazi

Late Rupelian

 Italy

Monteviale

D. darwini from Messel pit, Hesse, Germany, 48 million years old
Skull of D. hantoniensis
Diplocynodon cf. ratelii

D. darwini

Lutetian

 Germany

Messel pit

All specimens are from Messel pit of Germany. Synonyms are: D. ebertsi and D. hallense.

D. elavericus[3]

Middle Priabonian

 France

Domérat

All specimens came from Allier, Massif Central of France.

D. gervaisi

Earliest Rupelian

 France

Ronzon

Synonyms are: Saurocainus gervaisi.

D. hantoniensis

Early Priabonian

 United Kingdom

Headon Hill Formation

All specimens came from Hordwell, southern England. D. cf. hantoniensis is known from the Oligocene of Dordogne, France.

D. muelleri[4]

Middle Rupelian

 Spain

El Talladell

More than 100 are known, all from Lleida Province, Catalonia. Synonyms are: Hispanochampsa muelleri, D. guerini and D. marini.

D. ratelii

 France

Saint-Gérand-le-Puy*

D. ratelii is the type species of Diplocynodon. Most of the specimens came from Allier, Massif Central of France. Synonyms are: D. gracile.

D. tormis

Late Bartonian

 Spain

Salamanca

D. ungeri[2]

Middle Miocene

Synonyms are: Enneodon ungeri, D. steineri, and D. styriacus (see text).

*Locality and/or horizon of the type specimen.

Phylogeny

Cladogram based on Martin, 2010:[3]

Alligatoroidea

Leidyosuchus

Globidonta

Diplocynodontidae

Baryphracta

Diplocynodon

D. darwini

D. hantoniensis

D. ratelii

D. elavericus

D. muelleri

D. tormis

References

  1. Massimo Delfino and Thierry Smith (2012). "Reappraisal of the morphology and phylogenetic relationships of the middle Eocene alligatoroid Diplocynodon deponiae (Frey, Laemmert, and Riess, 1987) based on a three-dimensional specimen". Journal of Vertebrate Paleontology. 32 (6): 1358–1369. doi:10.1080/02724634.2012.699484.CS1 maint: uses authors parameter (link)
  2. Jeremy E. Martin and Martin Gross (2011). "Taxonomic clarification of Diplocynodon Pomel, 1847 (Crocodilia) from the Miocene of Styria, Austria". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 261 (2): 177–193. doi:10.1127/0077-7749/2011/0159.CS1 maint: uses authors parameter (link)
  3. Jeremy E. Martin (2010). "A new species of Diplocynodon (Crocodylia, Alligatoroidea) from the Late Eocene of the Massif Central, France, and the evolution of the genus in the climatic context of the Late Palaeogene". Geological Magazine. 147 (4): 596–610. doi:10.1017/S0016756809990161.CS1 maint: uses authors parameter (link)
  4. Paolo Pirasa and Angela D. Buscalionib (2006). "Diplocynodon muelleri comb. nov., an Oligocene diplocynodontine alligatoroid from Catalonia (Ebro Basin, Lleida Province, Spain)". Journal of Vertebrate Paleontology. 26 (3): 608–620. doi:10.1671/0272-4634(2006)26[608:DMCNAO]2.0.CO;2.CS1 maint: uses authors parameter (link)
  • Fossils (Smithsonian Handbooks) by David Ward (Page 243)


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