Eutyrannosauria

Eutyrannosaurians; Temporal range: ; Late Cretaceous, 80.6–66 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	Dryptosaurus aquilunguis
	Tyrannosaurus rex
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	Saurischia
Clade:	Theropoda
Superfamily:	†Tyrannosauroidea
Clade:	†Pantyrannosauria
Clade:	†Eutyrannosauria; Delcourt & Grillo, 2018
Subgroups
	†Dryptosaurus; †Appalachiosaurus; †Bistahieversor; †Tyrannosauridae;

Eutyrannosauria is a clade of tyrannosauroid theropods whose distribution has been found in what is now Asia and North America.[1] The clade consists of an evolutionary grade of tyrannosaurs such as Appalachiosaurus, Dryptosaurus, and Bistahieversor which led up to the family Tyrannosauridae.[2][3][1] The group was named in 2018 by Delcourt and Grillo in their paper about possible southern hemisphere tyrannosauroids and the phylogeography of tyrannosaurs.[1]

Classification

Below is a phylogeographic cladogram of Eutyrannosauria after Voris et al. (2020):[4]

Eutyrannosauria

Dryptosaurus aquilunguis

Appalachiosaurus montgomeriensis

Bistahieversor sealeyi

Tyrannosauridae

Albertosaurinae

	Gorgosaurus libratus

	Albertosaurus sarcophagus

Tyrannosaurinae

Alioramini

Qianzhousaurus sinensis

	Alioramus remotus

	Alioramus altai

	Teratophoneus curriei

	Dynamoterror dynastes

	Lythronax argestes

Nanuqsaurus hoglundi

Daspletosaurini

Thanatotheristes degrootorum

	Daspletosaurus torosus

	Daspletosaurus horneri

Tyrannosaurini

Zhuchengtyrannus magnus

	Tarbosaurus bataar

	Tyrannosaurus rex

References

Delcourt, R.; Grillo, O. N. (2018). "Tyrannosauroids from the Southern Hemisphere: Implications for biogeography, evolution, and taxonomy". Palaeogeography, Palaeoclimatology, Palaeoecology. 511: 379–387. Bibcode:2018PPP...511..379D. doi:10.1016/j.palaeo.2018.09.003.
Loewen, M.A.; Irmis, R.B.; Sertich, J.J.W.; Currie, P. J.; Sampson, S. D. (2013). Evans, David C (ed.). "Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans". PLoS ONE. 8 (11): e79420. Bibcode:2013PLoSO...879420L. doi:10.1371/journal.pone.0079420. PMC 3819173. PMID 24223179.
Stephen L. Brusatte, Alexander Averianov, Hans-Dieter Sues, Amy Muir and Ian B. Butler (2016). "New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs". Proceedings of the National Academy of Sciences of the United States of America. 113 (13): 3447–3452. doi:10.1073/pnas.1600140113. PMC 4822578. PMID 26976562.CS1 maint: multiple names: authors list (link)
Jared T. Voris; François Therrien; Darla K. Zelenitsky; Caleb M. Brown (2020). "A new tyrannosaurine (Theropoda:Tyrannosauridae) from the Campanian Foremost Formation of Alberta, Canada, provides insight into the evolution and biogeography of tyrannosaurids". Cretaceous Research. in press: Article 104388. doi:10.1016/j.cretres.2020.104388.

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[SouthernTyrannosaur-1] Delcourt, R.; Grillo, O. N. (2018). "Tyrannosauroids from the Southern Hemisphere: Implications for biogeography, evolution, and taxonomy". Palaeogeography, Palaeoclimatology, Palaeoecology. 511: 379–387. Bibcode:2018PPP...511..379D. doi:10.1016/j.palaeo.2018.09.003.

[Loewen13-2] Loewen, M.A.; Irmis, R.B.; Sertich, J.J.W.; Currie, P. J.; Sampson, S. D. (2013). Evans, David C (ed.). "Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans". PLoS ONE. 8 (11): e79420. Bibcode:2013PLoSO...879420L. doi:10.1371/journal.pone.0079420. PMC 3819173. PMID 24223179.

[Brusatte2016-3] Stephen L. Brusatte, Alexander Averianov, Hans-Dieter Sues, Amy Muir and Ian B. Butler (2016). "New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs". Proceedings of the National Academy of Sciences of the United States of America. 113 (13): 3447–3452. doi:10.1073/pnas.1600140113. PMC 4822578. PMID 26976562.CS1 maint: multiple names: authors list (link)

[Vorisetal2020-4] Jared T. Voris; François Therrien; Darla K. Zelenitsky; Caleb M. Brown (2020). "A new tyrannosaurine (Theropoda:Tyrannosauridae) from the Campanian Foremost Formation of Alberta, Canada, provides insight into the evolution and biogeography of tyrannosaurids". Cretaceous Research. in press: Article 104388. doi:10.1016/j.cretres.2020.104388.