List of procyonids
Procyonidae is a family of mammals in the order Carnivora, which includes raccoons, coatis, olingos, kinkajous, ring-tailed cats, and cacomistles, and many other extant and extinct mammals. A member of this family is called a procyonid. They are native to North and South America, though the common raccoon has been introduced to Europe, western Asia, and Japan. Procyonid habitats are generally forests, though some are found in shrublands and grasslands as well. The ring-tailed cat has a varied range including rocky areas and deserts as well as forests, and the common raccoon is widespread in urban environments. Species range in size from around 30–65 cm (12–26 in) long, plus a tail generally as long again. Population sizes are largely unknown, though the Cozumel raccoon is critically endangered, with around 200 individuals left, and the Eastern mountain coati is endangered. No procyonid species have been domesticated, although raccoons are sometimes kept as pets.
The fourteen species of Procyonidae are split into six genera, which are not currently grouped into named clades. Procyonidae is believed to have diverged as a separate family within Carnivora around 22.6 million years ago.[1] In addition to the extant species, as of 2020 Procyonidae includes forty extinct species placed in the six extant and nineteen extinct genera, though due to ongoing research and discoveries the exact number and categorization is not fixed.
Conventions
IUCN Red List categories | |
---|---|
Conservation status | |
EX | Extinct (0 species) |
EW | Extinct in the wild (0 species) |
CR | Critically endangered (1 species) |
EN | Endangered (1 species) |
VU | Vulnerable (0 species) |
NT | Near threatened (2 species) |
LC | Least concern (10 species) |
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the procyonid's range is provided. Ranges are based on the IUCN red list for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol (). Population figures are rounded to the nearest hundred.
Classification
The family Procyonidae consists of fourteen extant species belonging to six genera and divided into dozens of extant subspecies. This does not include hybrid species or extinct prehistoric species. Some prior classification schemes included the red panda or divided the family into named subfamilies and tribes based on similarities in morphology, though modern molecular studies indicate instead that the kinkajou is basal to the family, while raccoons, cacomistles, and ring-tailed cats form one clade and coatis and olingos another, despite morphology suggesting otherwise.
- Genus Bassaricyon (olingos): four species
- Genus Bassariscus (ring-tailed cats and cacomistles): two species
- Genus Nasua (coatis): two species
- Genus Nasuella (mountain coatis): two species
- Genus Potos (kinkajous): one species
- Genus Procyon (raccoons): three species
|
Procyonids
The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis; this includes rearranging Bassaricyon from five species to a mostly different four, and promoting the eastern mountain coati from a subspecies of the mountain coati.[2][3] There are additional proposals which are disputed, such as promoting the Guadeloupe raccoon population of the Bahamian raccoon subspecies of raccoon to a separate subspecies,[4] which are not included here.
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Eastern lowland olingo
|
B. alleni Thomas, 1880 |
Northwest South America |
Size: 30–46 cm (12–18 in) long, plus 40–53 cm (16–21 in) tail[2] Habitat: Forest[5] Diet: Primarily eats fruit, as well as small rodents, lizards, birds, insects, and eggs[5] |
LC
|
Northern olingo
|
B. gabbii Allen, 1876 |
Central America |
Size: 38–45 cm (15–18 in) long, plus 40–53 cm (16–21 in) tail[2] Habitat: Forest[6] Diet: Primarily eats fruit, nectar, flowers, insects, and small vertebrates[7][6] |
LC
|
Olinguito
|
B. neblina Helgen, 2013 Four subspecies
|
Andes mountains in northwest South America |
Size: 32–40 cm (13–16 in) long, plus 33–43 cm (13–17 in) tail[2] Habitat: Forest[8] Diet: Primarily eats fruit[8] |
NT
|
Western lowland olingo
|
B. medius Thomas, 1909 Two subspecies
|
Northwest South America and eastern Central America |
Size: 33–39 cm (13–15 in) long, plus 35–52 cm (14–20 in) tail[2] Habitat: Forest[9] Diet: Primarily eats fruit and nectar[9] |
LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Cacomistle
|
B. sumichrasti Saussure, 1860 Five subspecies
|
Southern Mexico and Central America |
Size: 38–47 cm (15–19 in) long, plus 39–53 cm (15–21 in) tail[10] Habitat: Forest[11] Diet: Primarily eats fruit, insects, and small vertebrates[11] |
LC
|
Ring-tailed cat
|
B. astutus Lichtenstein, 1830 Fourteen subspecies
|
Mexico and southwestern United States |
Size: 30–42 cm (12–17 in) long, plus 31–45 cm (12–18 in) tail[12] Habitat: Shrubland, forest, rocky areas, desert, and grassland[13] Diet: Primarily eats rodents, insects, birds, and fruit[13] |
LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
South American coati
|
N. nasua Linnaeus, 1766 Thirteen subspecies
|
Northern and central South America |
Size: 41–67 cm (16–26 in) long, plus 32–69 cm (13–27 in) tail[14] Habitat: Forest and shrubland[15] Diet: Primarily eats invertebrates and fruit[15] |
LC
|
White-nosed coati
|
N. narica Linnaeus, 1766 Four subspecies
|
Southern North America, Central America, and northwest South America |
Size: 33–67 cm (13–26 in) long, plus 33–67 cm (13–26 in) tail[16] Habitat: Grassland and forest[17] Diet: Primarily eats fruit and invertebrates[17] |
LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Eastern mountain coati | N. meridensis Thomas, 1901 |
Andes mountains in Venezuela |
Size: 43–54 cm (17–21 in) long, plus 19–30 cm (7–12 in) tail[3] Habitat: Forest and grassland[18] Diet: Unknown[18] |
EN
|
Western mountain coati
|
N. olivacea Gray, 1865 Two subspecies
|
Andes mountains in northern South America |
Size: 36–39 cm (14–15 in) long, plus 20–24 cm (8–9 in) tail[19] Habitat: Forest and grassland[20] Diet: Primarily eats invertebrates, small vertebrates, fruit, and vegetable remains[20] |
NT
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Kinkajou
|
P. flavus Schreber, 1774 Seven subspecies
|
Central America and northern South America |
Size: 40–60 cm (16–24 in) long, plus 40–60 cm (16–24 in) tail[21] Habitat: Forest[22] Diet: Primarily eats fruit, as well as flowers and leaves[22] |
LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Cozumel raccoon
|
P. pygmaeus Merriam, 1901 |
Cozumel island in Mexico |
Size: 42–60 cm (17–24 in) long, plus 23–26 cm (9–10 in) tail[23] Habitat: Forest[24] Diet: Primarily eats crabs, as well as fruit, insects, crayfish, and small vertebrates[24] |
CR
|
Crab-eating raccoon
|
P. cancrivorus G. Cuvier, 1798 Four subspecies
|
South America |
Size: 54–65 cm (21–26 in) long, plus 25–38 cm (10–15 in) tail[25] Habitat: Forest and inland wetlands[26] Diet: Primarily eats molluscs, fish, crabs, insects, and amphibians[26] |
LC
|
Raccoon
|
P. lotor Linnaeus, 1758 22 subspecies
|
North and Central America, and introduced to Central Europe, the Caucasus Mountains, and Japan |
Size: 41–55 cm (16–22 in) long, plus 19–41 cm (7–16 in) tail[27] Habitat: Forest[28] Diet: Omnivorous, eats fruit, nuts, insects, small mammals, eggs, birds, frogs, fish, aquatic invertebrates, worms, and garbage[28] |
LC
|
Prehistoric procyonids
In addition to extant procyonids, some prehistoric species have been discovered and classified as a part of Procyonidae. There is no generally accepted classification of extinct procyonid species. The species listed here are based on data from the Paleobiology Database, unless otherwise cited. Where available, the approximate time period the species was extant is given in millions of years before the present (Mya), also based on data from the Paleobiology Database.[29] All listed species are extinct; where a genus or subfamily within Procyonidae comprises only extinct species, it is indicated with a dagger symbol .
- Genus Angustictis (23–20 Mya)
- A. mayri (23–20 Mya)
- Genus Amphinasua
- Genus Arctonasua (16–4.9 Mya)
- A. eurybates (11–4.9 Mya)
- A. floridana (14–10 Mya)
- A. fricki (11–4.9 Mya)
- A. gracilis (16–13 Mya)
- A. minima (16–13 Mya)
- Genus Bassaricynoides (21–15 Mya)
- B. phyllismillerae (21–15 Mya)
- B. stewartae (21–15 Mya)
- Genus Brachynasua
- B. merani
- Genus Broiliana
- B. dehmi
- B. nobilis (16–13 Mya)
- Genus Chapalmalania
- C. altaefrontis
- C. ortognatha
- Genus Cyonasua
- C. argentina (9.0–6.8 Mya)
- C. brevirrostris
- C. clausa (6.8–4.0 Mya)
- C. groeberi (9.0–4.0 Mya)
- C. longirostris
- C. lutaria
- C. meranii
- C. pascuali (9.0–6.8 Mya)
- C. robusta
- Genus Edaphocyon (21–13 Mya)
- E. lautus (21–15 Mya)
- E. palmeri (21–15 Mya)
- E. pointblankensis (16–13 Mya)
- Genus Myxophagus
- M. spelaeus
- Genus Nasua (10 Mya–present)[2]
- N. nicaeensis
- N. pronarica (4.9–1.8 Mya)
- Genus Pachynasua
- P. clausa
- Genus Parahyaenodon (6.8–4.0 Mya)
- P. argentinus (6.8–4.0 Mya)
- Genus Paranasua (14–10 Mya)
- P. biradica (14–10 Mya)
- Genus Parapotos (16–13 Mya)
- P. tedfordi (16–13 Mya)
- Genus Probassariscus (16–13 Mya)
- P. matthewi (16–13 Mya)
- Genus Procyon (10 Mya–present)[2]
- P. rexroadensis (4.9–1.8 Mya)
- Genus Protoprocyon (14–10 Mya)
- P. savagei (14–10 Mya)
- Genus Sivanasua
- S. viverroides
- Genus Stromeriella (23–13 Mya)
- S. depressa (23–20 Mya)
- S. franconica (16–13 Mya)
- Genus Tetraprothomo (6.8–4.0 Mya)
- T. argentinus (6.8–4.0 Mya)
References
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