2011 in paleontology

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2011.

List of years in paleontology (table)
In science
2008
2009
2010
2011
2012
2013
2014

Protozoa

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Vetufebrus[2]

Gen et sp nov

Valid

Poinar

Burdigalian

Dominican amber

 Dominican Republic

A Plasmodiidae vectored by the bat fly Enischnomyia stegosoma

Plants

Angiosperms

Name Novelty Status Authors Age Unit Location Notes Images

Palmoxylon lametaei[3]

Species

Valid

Dutta, Ambwani, & Estrada-Ruiz

Maastrichtian

Lameta Formation

 India

A petrified palm wood

Leefructus[4]

Gen. et sp.

Valid

Sun, Dilcher, Wang, & Chen

Early Cretaceous

Yixian Formation

 China

A basal eudicot

Gymnosperms

Name Novelty Status Authors Age Unit Location Notes Images

Qataniaria[5]

Gen. et sp.

Valid

Krassilov & Schrank

Albian

Hatira Formation

 Israel

Cariria[6]

Gen. et sp.

Valid

Kunzmann, Mohr, Wilde & Bernardes-de-Oliveira

Aptian

Crato Formation

 Brazil

Pteridophytes

Name Novelty Status Authors Age Unit Location Notes Images

Equisetum thermale[7]

sp. nov

Valid

Channing, Zamuner, Edwards, & Guido

Callovian - Tithonian

La Matilde Formation

 Argentina

oldest crown group Equisetum species

Nematoda

Name Novelty Status Authors Age Unit Location Notes Images

Formicodiplogaster[8]

Gen et sp nov

valid

Poinar

Burdigalian

Dominican amber

 Dominican Republic

Diplogasteridae nematode, parasite of ant genus Azteca;
Type species Formicodiplogaster myrmenema

Lobopods

Name Novelty Status Authors Age Unit Location Notes Images

Diania cactiformis[9]

Valid

Liu et al.

Lower Cambrian

Maotianshan shale

 China

A lobopodian.

Mureropodia apae

Valid

Gamez Vintaned, Linan, & Zhuravlev

Early Cambrian

Murero

 Spain

A lobopodian.

Siberion lenaicus

Valid

Dzik

Early Cambrian

Sinsk Formation

 Russia

A lobopodian.

Vetulicolians

Name Novelty Status Authors Age Unit Location Notes Images

Ooedigera peeli[10]

gen. nov

valid

Zinther, et al.

Early Cambrian

Sirius Passet

 Greenland

earliest known vetulicolian described

Artist's reconstruction

Molluscs

Arthropods

Fishes

Amphibians

Newly named lepospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Kirktonecta[11]

Gen. et sp.

Valid

  • Clack

Viséan

East Kirkton Quarry

A microsaur.

Tambaroter[12]

Gen. et sp.

Valid

Early Permian

Tambach Formation

An ostodolepid.

Newly named temnospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Acheloma dunni[13]

Species

Valid

  • Polley
  • Reisz

Early Permian

A trematopid.

Calmasuchus[14]

Gen. et sp.

Valid

  • Fortuny
  • Galobart
  • De Santisteban

early-middle Anisian

Catalan basin

A capitosaurian.

Rotaryus[15]

Gen. et sp.

Valid

Early Permian

Tambach Formation

A trematopid.

Rubeostratilia[16]

Gen. et sp.

Valid

  • Bourget
  • Anderson

Early Permian

An amphibamid.

Rubeostratilia texensis.

Newly named lissamphibians

Name Novelty Status Authors Age Unit Location Notes Images

Urupia[17]

Gen. et sp.

Valid

  • Skutschas
  • Krasnolutskii

Bathonian

Itat Formation

A basal stem salamander.

Basal reptiles

Newly named captorhinids

Name Novelty Status Authors Age Unit Location Notes Images

Gansurhinus[18]

Gen. et sp.

Valid

Middle Permian

A moradisaurine captorinid.

Newly named basal diapsids

Name Novelty Status Authors Age Unit Location Notes Images

Orovenator[19]

Gen. et sp.

Valid

Sakmarian

The oldest and most basal neodiapsid.

Newly named ichthyosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Sveltonectes[20]

Gen. et sp.

Valid

Barremian

An ophthalmosaurid.

Lepidosauromorphs

Newly named saurosphargids

Name Novelty Status Authors Age Unit Location Notes Images

Sinosaurosphargis[21]

Gen. et sp.

Valid

Middle Anisian

Guanling Formation

An incertae sedis.

Newly named sauropterygians

Name Novelty Status Authors Age Unit Location Notes Images

Abyssosaurus[22]

Gen. et sp.

Valid

  • Berezin

Late Hauterivian

An aristonectid.

Diandongosaurus[23]

Gen. et sp.

Valid

  • Shang
  • Wu
  • Li

Middle Anisian

Guanling Formation

A relative of nothosauroids.[24]

Dianopachysaurus[25]

Gen. et sp.

Valid

Middle Anisian

Guanling Formation

A keichousaurid pachypleurosaur.

Hauffiosaurus tomistomimus[26]

Species

Valid

Early Toarcian

Whitby Limestone Formation

A second species of Hauffiosaurus.

Marmornectes[27]

Gen. et sp.

Valid

  • Ketchum
  • Benson

Callovian

Oxford Clay Formation

A pliosaurid.

Westphaliasaurus[28]

Gen. et sp.

Valid

  • Schwermann
  • Sander

Pliensbachian

Höxter district

A plesiosaurid plesiosaur.

Zarafasaura[29]

Gen. et sp.

Valid

Maastrichtian

An elasmosaurid plesiosaur.

Newly named lizards

Name Novelty Status Authors Age Unit Location Notes Images

Brasiliguana[30]

Gen. et sp.

Valid

  • Nava
  • Martinelli

Turonian/Santonian

Adamantina Formation

An iguanian lizard.

Cryptolacerta[31]

Gen. et sp.

Valid

Lutetian

Messel pit

An amphisbaenian lizard.

Latoplatecarpus[32]

Gen. et sp.

Valid

  • Konishi
  • Caldwell

Campanian

Pierre Shale Formation
Demopolis Chalk Formation

A plioplatecarpine mosasaur. Its type species is Latoplatecarpus willistoni; "Plioplatecarpus" nichollsae (Cuthbertson et al.., 2007) is a second species of Latoplatecarpus.

Oreithyia[33][34]

Gen. et sp. nov

Valid

  • Smith

Eocene (Chadronian)

A member of Corytophanidae. The type species is Oreithyia oaklandi.

Plesioplatecarpus[32]

Gen. et comb.

Valid

  • Konishi
  • Caldwell

Middle Coniacian to middle Santonian.

Niobrara Chalk
Eutaw Formation

A plioplatecarpine mosasaur, a new genus for "Clidastes" planifrons (Cope, 1874).

Queironius[33]

Gen. et sp. nov

Valid

  • Smith

Eocene (Chadronian)

A relative of the desert iguana. The type species is Queironius praelapsus.

Sauropithecoides[33]

Gen. et comb. nov

Valid

  • Smith

Eocene (Chadronian)

A member of Polychrotidae; a new genus for "Polychrus" charisticus Smith (2006).

Newly named snakes

Name Novelty Status Authors Age Unit Location Notes Images

Madtsoia pisdurensis[35]

Species

Valid

  • Mohabey
  • Head
  • Wilson

Late Cretaceous

A madtsoiid snake.

Turtles

Newly named turtles

Name Novelty Status Authors Age Unit Location Notes Images

Albertwoodemys[36]

Gen. et sp.

Valid

Early Oligocene

Jebel Qatrani Formation

A podocnemidid.

Brontochelys[36]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Early Miocene

A podocnemidid, a new genus for "Shweboemys" gaffneyi.

Cordichelys[36]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Eocene

Qasr el-Sagha beds

A podocnemidid, a new genus for "Podocnemis" antiqua.

Gilmoremys[37]

Gen. et comb.

Valid

  • Joyce
  • Lyson

Maastrichtian

Lance Formation
Hell Creek Formation

A softshell turtle, new genus for "Aspideretes" lancensis (Gilmore, 1928).

Graptemys kerneri[38]

Sp. nov

Valid

  • Ehret & Bourque

Late Pleistocene

An emydid, a species of Graptemys.

Hispaniachelys[39]

Gen. et sp.

Valid

Oxfordian

Lorente Formation

A paracryptodire or a basal testudine.

Hoplochelys clark[40]

Species

Valid

Maastrichtian

Hell Creek Formation

A kinosternoid, a species of Hoplochelys.

Lapparentemys[36]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Paleocene

Santa Lucía Formation

A podocnemidid, a new genus for "Roxochelys" vilavilensis.

Latentemys[36]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Uncertain, possibly Miocene.

Uncertain, possibly Moghara Formation.

A podocnemidid.

Lemurchelys[36]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Early Miocene

Moghara Formation

A podocnemidid.

Mogharemys[36]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Miocene

A podocnemidid, a new genus for "Sternothaerus" blanckenhorni.

Oliveremys[41]

Gen. et comb. nov.

Valid

  • Vitek

Eocene

Washakie Formation

A trionychid, a new genus for "Trionyx" uintaensis.

Patagoniaemys[42]

Gen. et sp.

Valid

  • Sterli
  • De la Fuente

Campanian - Maastrichtian

La Colonia Formation

A stem turtle.

Peiropemys[36]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Cretaceous

Marília Formation

A podocnemidid.

Pricemys[36]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Cretaceous

Marília Formation

A podocnemidid.

Prochelidella cerrobarcinae[43]

Species

Valid

Aptian/Albian

Cerro Barcino Formation

The oldest known chelid, a species of Prochelidella.

Selenemys[44]

Gen. et sp.

Valid

  • Pérez-García
  • Ortega

Upper Kimmeridgian

Lusitanian Basin

A pleurosternid.

Sokatra[45]

Gen. et sp.

Valid

  • Gaffney
  • Krause

Maastrichtian

Maevarano Formation

A pleurodiran.

Trionyx ikoviensis[46]

Species

Valid

Middle Eocene

A species of Trionyx sensu lato.

Wilburemys[47]

Gen. et sp. nov

Valid

Late Miocene-Pliocene (Hemphillian-Blancan)

Ellensburg Formation

A member of Emydidae belonging to the subfamily Emydinae. The type species is Wilburemys yakimensis.

Archosauromorphs

Newly named basal archosauromorphs

Name Novelty Status Authors Age Unit Location Notes Images

Archeopelta[48]

Gen. et sp.

Valid

  • Desojo
  • Ezcurra
  • Schultz

Late Ladinian/Early Carnian

Santa Maria Formation

A member of Archosauriformes of uncertain phylogenetic placement. Originally classified as a doswelliid, but subsequently argued to be an erpetosuchid archosaur.[49]

Augustaburiania[50]

Gen. et sp.

Valid

  • Sennikov

Early Triassic

Don River Basin

The earliest tanystropheid.

Kyrgyzsaurus[51]

Gen. et sp.

Valid

  • Alifanov
  • Kurochkin

Late Ladinian/Early Carnian

Madygen Formation

The earliest drepanosaurid.

Protanystropheus[50]

Gen. et comb.

Valid

  • Sennikov

Middle - Late Triassic

A tanystropheid, a new genus for "Tanystropheus" antiquus (von Huene, 1905).

Archosaurs

Synapsids

Newly named non-mammalian synapsids

Name Novelty Status Authors Age Unit Location Notes Images

Annatherapsidus postum[52]

Sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Basilodon[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" woodwardi.

Bridetherium[54]

Gen. et sp.

Valid

  • Clemens

Early Jurassic

A morganucodontan.

Candelariodon[55]

Gen. et sp.

Valid

Middle Triassic

Santa Maria Formation

A carnivorous cynodont.

Cryptovenator[56]

Gen. et sp.

Valid

Late Gzhelian

Remigiusberg Formation

A sphenacodontid pelycosaur. The type species is Cryptovenator hirschbergeri.

Euptychognathus[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" bathyrhynchus.

Euromycter[57]

Gen. et comb.

Valid

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

A large caseid. A new genus for "Casea" rutena.

Gorochovetzia[52]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Keyseria[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" benjamini.

Muchia[52]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Paceyodon[54]

Gen. et sp.

Valid

  • Clemens

Early Jurassic

A morganucodontan.

Peramodon[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" amalitzkii

Perplexisaurus lepusculus[52]

Species

Valid

  • Ivakhnenko

Middle Permian

A therocephalian.

Purlovia[52]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian with a short skull.

Ruthenosaurus[57]

Gen. et sp.

Valid

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

A large caseid. The type species is Ruthenosaurus russellorum

Syops[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" vanhoepeni.

Tiarajudens[58]

Gen. et sp.

Valid

Capitanian

Rio do Rasto Formation

A basal, saber-toothed herbivorous anomodont. The type species is Tiarajudens eccentricus.

Mammals

Footnotes

Complete author list

As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.

  1. Henrici, Martens, Berman, Sumida.
  2. Berman, Henrici, Martens, Sumida, Anderson
  3. Reisz, Liu, Li, Müller.
  4. Fischer, Masure, Arkhangelsky, Godefroit.
  5. Li, Rieppel, Wu, Zhao, Wang.
  6. Liu, Rieppel, Jiang, Aitchison, Motani, Zhang, Zhou, Sun.
  7. Benson, Ketchum, Noè, Gómez-Pérez.
  8. Vincent, Bardet, Suberbiola, Bouya, Amaghzaz, Meslouh.
  9. Müller, Hipsley, Head, Kardjilov, Hilger, Wuttke, Reisz.
  10. Gaffney, Meylan, Wood, Simons and Campos
  11. Slater, Reolid, Wood, Schouten and Benton
  12. Knauss, Joyce, Lyson and Pearson
  13. de la Fuente, Umazano, Sterli and Carballido
  14. Danilov, Zvonok, Syromyatnikova and Udovichenko
  15. De Oliveira, Schultz, Soares, Rodrigues.
  16. Fröbisch, Schoch, Müller, Schindler, Schweiss.
  17. Reisz, Maddin, Fröbisch, Falconnet.
  18. Cisneros, Abdala, Rubidge, Dentzien-Dias, Bueno.

References

  1. Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
  2. Poinar, G. (2011). "Vetufebrus ovatus n. gen., n. sp. (Haemospororida: Plasmodiidae) vectored by a streblid bat fly (Diptera: Streblidae) in Dominican amber". Parasites & Vectors. 4 (229): 1–5. doi:10.1186/1756-3305-4-229. PMC 3253689. PMID 22152687.
  3. Dutta, D.; Ambwani, K.; Estrada-Ruiz, E. (2011). "Late Cretaceous palm stem Palmoxylon lametaei sp. nov. from Bhisi Village, Maharashtra, India" (PDF). Revista Mexicana de Ciencias Geológicas. 28 (1): 1–9.
  4. Ge Sun; David L. Dilcher; Hongshan Wang; Zhiduan Chen (2011). "A eudicot from the Early Cretaceous of China". Nature. 471 (7340): 625–628. Bibcode:2011Natur.471..625S. doi:10.1038/nature09811. PMID 21455178. S2CID 205224036.
  5. Valentin Krassilov; Eckart Schrank (2011). "New Albian macro- and palynoflora from the Negev (Israel) with description of a new gymnosperm morphotaxon" (PDF). Cretaceous Research. 32 (1): 13–29. doi:10.1016/j.cretres.2010.10.001. Archived from the original (PDF) on 2012-03-30. Retrieved 2011-08-20.
  6. Kunzmann, L.; Mohr, B.A.R.; Wilde, V.; Bernardes-de-Oliveira, M.E.C. (2011). "A putative gnetalean gymnosperm Cariria orbiculiconiformis gen. nov. et spec. nov. from the Early Cretaceous of northern Gondwana". Review of Palaeobotany and Palynology. 165 (1–2): 75–95. doi:10.1016/j.revpalbo.2011.02.005.
  7. Channing, A.; Zamuner, A.; Edwards, D.; Guido, D. (2011). "Equisetum thermale sp. nov. (Equisetales) from the Jurassic San Agustin hot spring deposit, Patagonia: Anatomy, paleoecology, and inferred paleoecophysiology". American Journal of Botany. 98 (4): 680–697. doi:10.3732/ajb.1000211. PMID 21613167.
  8. Poinar, G.; Heiss, E. (2011). "New Termitaphididae and Aradidae (Hemiptera) in Mexican and Dominican amber" (PDF). Palaeodiversity. 4: 51–62.
  9. Liu, J.; Steiner, M.; Dunlop, J. A.; Keupp, H.; Shu, D.; Ou, Q.; Han, J.; Zhang, Z.; Zhang, X. (2011). "An armoured Cambrian lobopodian from China with arthropod-like appendages". Nature. 470 (7335): 526–530. Bibcode:2011Natur.470..526L. doi:10.1038/nature09704. PMID 21350485. S2CID 4324509.
  10. Vinther, J.; Smith, M. P.; Harper, D. A. T. (2011). "Vetulicolians from the Lower Cambrian Sirius Passet Lagerstätte, North Greenland, and the polarity of morphological characters in basal deuterostomes". Palaeontology. 54 (3): 711–719. doi:10.1111/j.1475-4983.2011.01034.x.
  11. Clack, Jennifer A. (2011). "A new microsaur from the early Carboniferous (Viséan) of East Kirkton, Scotland, showing soft tissue evidence". Special Papers in Palaeontology. 86: 45–55. doi:10.1111/j.1475-4983.2011.01073.x (inactive 2021-01-14).CS1 maint: DOI inactive as of January 2021 (link)
  12. Henrici, A.C.; Martens, T.; Berman, D.S.; Sumida, S.S. (2011). "An ostodolepid 'microsaur' (Lepospondyli) from the Lower Permian Tambach Formation of central Germany". Journal of Vertebrate Paleontology. 31 (5): 997–1004. doi:10.1080/02724634.2011.596601. S2CID 129710688.
  13. Polley, Brendan P.; Reisz, Robert R. (2011). "A new Lower Permian trematopid (Temnospondyli: Dissorophoidea) from Richards Spur, Oklahoma". Zoological Journal of the Linnean Society. 161 (4): 789–815. doi:10.1111/j.1096-3642.2010.00668.x.
  14. Fortuny, Josep; Galobart, Àngel; De Santisteban, Carles (2011). "A new capitosaur from the Middle Triassic of Spain and the relationships within the Capitosauria" (PDF). Acta Palaeontologica Polonica. 56 (3): 553–66. doi:10.4202/app.2010.0025. S2CID 55068128. Archived from the original (PDF) on 2011-05-22.
  15. David S. Berman; Amy C. Henrici; Thomas Martens; Stuart S. Sumida; Jason S. Anderson (2011). "Rotaryus gothae, a New Trematopid (Temnospondyli: Dissorophoidea) from the Lower Permian of Central Germany". Annals of Carnegie Museum. 80 (1): 49–65. doi:10.2992/007.080.0106. S2CID 84780478.
  16. Bourget, Hélène; Anderson, Jason S. (2011). "A new amphibamid (Temnospondyli: Dissorophoidea) from the Early Permian of Texas". Journal of Vertebrate Paleontology. 31 (1): 32–49. doi:10.1080/02724634.2011.539652. S2CID 85856972.
  17. P.P. Skutschas; S.A. Krasnolutskii (2011). "A new genus and species of basal salamanders from the Middle Jurassic of western Siberia, Russia" (PDF). Proceedings of the Zoological Institute of the Russian Academy of Sciences. 315 (2): 167–175.
  18. Robert R. Reisz; Jun Liu; Jin-Ling Li; Johannes Müller (2011). "A new captorhinid reptile, Gansurhinus qingtoushanensis, gen. et sp. nov., from the Permian of China". Naturwissenschaften. 98 (5): 435–441. Bibcode:2011NW.....98..435R. doi:10.1007/s00114-011-0793-0. PMID 21484260. S2CID 20274349.
  19. Robert R. Reisz; Sean P. Modesto; Diane M. Scott (2011). "A new Early Permian reptile and its significance in early diapsid evolution". Proceedings of the Royal Society B. 278 (1725): 3731–3737. doi:10.1098/rspb.2011.0439. PMC 3203498. PMID 21525061.
  20. Fischer, V.; Masure, E.; Arkhangelsky, M.S.; Godefroit, P. (2011). "A new Barremian (Early Cretaceous) ichthyosaur from western Russia". Journal of Vertebrate Paleontology. 31 (5): 1010–1025. doi:10.1080/02724634.2011.595464. S2CID 86036325.
  21. Li, Chun; Olivier Rieppel; Xiao-Chun Wu; Li-Jun Zhao; Li-Ting Wang (2011). "A new Triassic marine reptile from southwestern China". Journal of Vertebrate Paleontology. 31 (2): 303–312. doi:10.1080/02724634.2011.550368. S2CID 130698779.
  22. A. Yu. Berezin (2011). "A new plesiosaur of the family Aristonectidae from the early cretaceous of the center of the Russian platform". Paleontological Journal. 45 (6): 648–660. doi:10.1134/S0031030111060037. S2CID 129045087.
  23. Shang Qing-Hua; Wu Xiao-Chun; Li Chun (2011). "A new eosauropterygian from Middle Triassic of eastern Yunnan Province, southwestern China" (PDF). Vertebrata PalAsiatica. 49 (2): 155–171. doi:10.19615/j.cnki.1000-3118.2011.02.002.
  24. Tamaki Sato; Yen-Nien Cheng; Xiao-Chun Wu; Hsi-Yin Shan (2013). "Diandongosaurus acutidentatus Shang, Wu & Li, 2011 (Diapsida: Sauropterygia) and the relationships of Chinese eosauropterygians". Geological Magazine. 151 (1): 121–133. Bibcode:2014GeoM..151..121S. doi:10.1017/S0016756813000356.
  25. Jun Liu; Olivier Rieppel; Da-Yong Jiang; Jonathan C. Aitchison; Ryosuke Motani; Qi-Yue Zhang; Chang-Yong Zhou; Yuan-Yuan Sun (2011). "A new pachypleurosaur (Reptilia: Sauropterygia) from the lower Middle Triassic of southwestern China and the phylogenetic relationships of Chinese pachypleurosaurs". Journal of Vertebrate Paleontology. 31 (2): 292–302. doi:10.1080/02724634.2011.550363. S2CID 86594206.
  26. Roger B. J. Benson; Hilary F. Ketchum; Leslie F. Noè; Marcela Gómez-Pérez (2011). "New information on Hauffiosaurus (Reptilia, Plesiosauria) based on a new species from the Alum Shale Member (Lower Toarcian: Lower Jurassic) of Yorkshire, UK" (PDF). Palaeontology. 54 (3): 547–571. doi:10.1111/j.1475-4983.2011.01044.x.
  27. Hilary F. Ketchum; Roger B. J. Benson (2011). "A new pliosaurid (Sauropterygia, Plesiosauria) from the Oxford Clay Formation (Middle Jurassic, Callovian) of England: evidence for a gracile, longirostrine grade of Early–Middle Jurassic pliosaurids". Special Papers in Palaeontology. 86: 109–129. doi:10.1111/j.1475-4983.2011.01083.x (inactive 2021-01-14).CS1 maint: DOI inactive as of January 2021 (link)
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