2019 in arthropod paleontology

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This list of fossil arthropods described in 2019 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that are scheduled to be described during the year 2019, as well as other significant discoveries and events related to arthropod paleontology that are scheduled to occur in the year 2019.

General research

A revision of higher-level arthropod taxonomy is published by Aria (2019), who proposes the name Cenocondyla for the least inclusive group containing both Mandibulata and Chelicerata.[1]
A study on the molecular composition and microanatomy of the eyes of 54-million-year-old crane flies from the Fur Formation (Denmark), evaluating its implications for the knowledge of optical systems of other ancient arthropods (especially trilobites), is published by Lindgren et al. (2019).[2]

Arachnids

Research

Two new specimens of laniatorid harvestmen (an adult specimen assignable to the species Proholoscotolemon nemastomoides and a smaller specimen belonging or related to the genus Proholoscotolemon) are described from the Eocene Baltic amber by Bartel & Dunlop (2019).[3]
A review of the phylogenetic hypotheses of scorpion interrelationships, as well as of scorpion palaeontology, is published by Howard et al. (2019), who also provide a list of scorpion fossil calibrations for use in molecular dating.[4]
A fossil spider, possibly a juvenile female golden silk orb-weaver, is described from the Lower Eocene Palana Formation of the Gurha opencast lignite mine (western Rajasthan, India) by Patel, Rana & Selden (2019), representing the first fossil spider from India which wasn't found in amber.[5]
A review of the fossil record and evolutionary history of spiders, reevaluating the phylogenetic placement of key fossils and aiming to determine whether there has been a major turnover in the spider fauna between the Mesozoic and Cenozoic, is published online by Magalhaes et al. (2019).[6]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Caestaranea[7]	Gen. et sp. nov	Valid	Selden, Huang & Garwood	Middle–Late Jurassic	Haifanggou Formation	China	A palpimanoid spider. Genus includes new species C. jurassica.
Colossopantopodus nanus[8]	Sp. nov	Valid	Sabroux et al.	Late Jurassic	Solnhofen Limestone	Germany	A sea spider.
Cosmobunus sagani[9]	Sp. nov	Valid	Palencia et al.	Miocene (Burdigalian)	Rubielos de Mora Basin	Spain	A member of Opiliones, a species of Cosmobunus.
Eatoniana crinita[10]	Sp. nov	Valid	Sidorchuk et al.	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A mite belonging to the family Erythraeidae.
Eomesothele[11]	Gen. et sp. nov	Valid	Wunderlich	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to group Mesothelae, assigned to the new family Eomesothelidae. The type species is E. noninclinata.
Eumesosoma abdelmawlai[12]	Sp. nov	Valid	Elsaka, Mitov & Dunlop	Eocene	Baltic amber	Europe (Baltic Sea region)	A harvestman, a species of Eumesosoma.
?Eurycyde golem[8]	Sp. nov	Valid	Sabroux et al.	Late Jurassic	Solnhofen Limestone	Germany	A sea spider belonging to the family Ascorhynchidae.
Intermesothele[11]	Gen. et sp. nov	Valid	Wunderlich	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A spider belonging to group Mesothelae, possibly belonging to the family Eomesothelidae. The type species is I. pulcher.
Jinjumegops[13]	Gen. et sp. nov	Valid	Park, Nam & Selden	Early Cretaceous (Albian)	Jinju Formation	South Korea	A spider belonging to the family Lagonomegopidae. The type species is J. dalingwateri.
Koreamegops[13]	Gen. et sp. nov	Valid	Park, Nam & Selden	Early Cretaceous (Albian)	Jinju Formation	South Korea	A spider belonging to the family Lagonomegopidae. The type species is K. samsiki.
Lutetiana[14]	Gen. et sp. nov	Valid	Selden & Wappler	Eocene (Lutetian)	Messel pit	Germany	A spider, possibly a member of the family Cybaeidae. The type species is L. neli.
Metagonia esquincacanoi[15]	Sp. nov		García-Villafuerte	Miocene (Aquitanian)	Mexican amber	Mexico	A species of Metagonia.
Neoliodes andreneli[16]	Sp. nov	Valid	Arillo et al.	Early Cretaceous	Lebanese amber	Lebanon	A mite belonging to the family Neoliodidae.
Onychopalpus[7]	Gen. et sp. nov	Valid	Selden, Huang & Garwood	Middle–Late Jurassic	Haifanggou Formation	China	A palpimanoid spider. Genus includes new species O. thomisoides.
Paragilarovia[12]	Gen. et sp. nov	Valid	Elsaka, Mitov & Dunlop	Eocene	Baltic amber	Europe (Baltic Sea region)	A harvestman belonging to the family Nemastomatidae. Genus includes new species P. hochae.
Procaeculus coineaui[17]	Sp. nov	Valid	Porta et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A mite belonging to the family Caeculidae.
Protolophus hoffeinsi[12]	Sp. nov	Valid	Elsaka, Mitov & Dunlop	Eocene	Baltic amber	Europe (Baltic Sea region)	A harvestman, a species of Protolophus.
Scolotydaeus vlaskini[18]	Sp. nov	Valid	Klimov et al.	Late Eocene	Rovno amber	Ukraine	A mite belonging to the family Paratydeidae.
Sinaranea brevicrus[7]	Sp. nov	Valid	Selden, Huang & Garwood	Middle–Late Jurassic	Haifanggou Formation	China	A palpimanoid spider.
Tanytydeus pogrebnyaki[18]	Sp. nov	Valid	Klimov et al.	Late Eocene	Rovno amber	Ukraine	A mite belonging to the family Paratydeidae.

Crustaceans

Research

A global dataset of exceptionally preserved muscles and muscle attachment scars in fossil malacostracan specimens is compiled by Klompmaker et al. (2019).[19]
A three-dimensionally preserved tanaidacean specimen belonging or related to the species Jurapseudes friedericianus is described from the Middle Jurassic (Callovian) Ornatenton Formation (Germany) by Schädel et al. (2019).[20]
A study on the evolutionary history of decapod crustaceans is published by Wolfe et al. (2019).[21]
A study on the chemical composition of two specimens of fossilized shrimps from the Cretaceous Romualdo and Ipubi formations (Araripe Basin, Brazil), and on its implications for inferring the course of the processes of fossilization of these crustaceans, is published by Barros et al. (2019).[22]
A study on the Middle Jurassic palaeoenvironment of La Voulte (France), as indicated by data from exceptionally preserved eyes of the polychelidan lobster Voulteryon parvulus and from epibiontic brachiopods associated with V. parvulus, is published by Audo et al. (2019).[23]
A study on the anatomy of the holotype specimen of a putative Paleocene shark Platyacrodus unicus is published by Bogan, Agnolin & Ezcurra (2019), who reinterpret this specimen as a carapace of a small retroplumid crab belonging to the genus Costacopluma.[24]
An achelatan larval fossil with an intermediate type of morphology, interpreted as the oldest occurrence of an achelatan lobster larva reported so far, is described from the Toarcian Posidonia Shale (Germany) by Haug, Haug & Schweigert (2019).[25]
Redescription of the isopod species Palaega sismondai Ristori (1891) is published by Hyžný, Pasini & Garassino (2019), who transfer this species to the genus Bathynomus.[26]
A moult of a giant isopod is described from the early Oligocene Ranzano Formation by Pasini & Garassino (2019), representing the oldest fossil record of the genus Bathynomus from Italy.[27]
A study on the Late Devonian ostracods from the Yangdi and Nandong sections in South China, and on their responses to the Frasnian-Famennian event, is published by Song, Huang & Gong (2019).[28]
A diverse earliest Triassic ostracod fauna is reported from the Yangou section in South China by Qiu et al. (2019).[29]
A revision of ostracods from the Upper Oligocene to Lower Miocene Pirabas Formation (Brazil) is published by Nogueira, Ramos & Hunt (2019).[30]
A study on changes of diversity of Neogene fossil marine ostracods from Java, Indonesia is published by Shin et al. (2019).[31]
A study on deep-sea benthic ostracod assemblages from the southern Sea of Japan covering the last 2 millions years, evaluating their responses to the Mid-Brunhes Event, orbital-scale climatic cycles, and fluctuations of the Tsushima Warm Current, is published by Huang et al. (2019).[32]
A three-dimensionally preserved specimen of a late growth stage of the phosphatocopid species Cyclotron angelini is described from the Furongian strata of northern Poland by Olempska et al. (2019).[33]
A study on the ecology of clam shrimps from the Anjiagou and Hengdaozi beds of the Lower Cretaceous Yixian Formation (China) is published by Hethke et al. (2019).[34]
A study on the oxygen isotope composition of whale barnacle shells from three Pleistocene localities along the eastern Pacific coast, and on their implications for the knowledge of the history of whale migrations, is published by Taylor et al. (2019).[35]
A study on the internal soft-tissue anatomy of Hesslandona angustata is published by Liu et al. (2019).[36]

Malacostracans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Acanthocarpus osborni[37]	Sp. nov	Valid	Feldmann, Schweitzer & Phillips	Oligocene		United States	A crab, a species of Acanthocarpus.
Ahazianassa[38]	Gen. et comb. nov	Valid	Karasawa et al.	Late Cretaceous (Campanian–Maastrichtian)	Izumi Group	Japan	A member of the family Gourretiidae; a new genus for "Callianassa" masanorii Karasawa (1998).
Alontecarcinus[39]	Gen. et sp. nov	Valid	De Angeli & Caporiondo	Eocene (Bartonian)		Italy	A crab belonging to the family Potamidae. The type species is A. buratoi.
Amatukamius[40]	Gen. et comb. nov	Valid	Karasawa & Ohara	Early Cretaceous (Barremian)	Arida Formation	Japan	A member of Axiidea belonging to the family Micheleidae; a new genus for "Callianassa" sakakuraorum Karasawa (2000).
Anapagurus dalessandroi[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Paguridae.
Annuntidiogenes elongatus[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Annuntidiogenidae.
Annuntidiogenes hoelderi[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Annuntidiogenidae.
Anoplocarcinus[43]	Gen. et sp. nov	Valid	Schweitzer et al.	Early Cretaceous (Albian)	Hudspeth Formation	United States	A crab belonging to the group Raninoida and to the family Orithopsidae. The type species is A. hudspethi.
Aragolambrus[44]	Gen. et sp. nov	Valid	Ferratges, Zamora & Aurell	Eocene (Ypresian)		Spain	A crab belonging to the family Parthenopidae. Genus includes new species A. collinsi.
Archaeochiapasa[45]	Gen. et sp. nov	Valid	Guinot, Carbot-Chanona & Vega	Late Cretaceous (Cenomanian)	Sierra Madre Formation	Mexico	A crab belonging to the group Heterotremata, assigned to the new family Archaeochiapasidae. The type species is A. mardoqueoi.
Archaeopus burgessi[46]	Sp. nov	Valid	Nyborg et al.	Late Cretaceous (Campanian)	Cedar District Formation	Canada	A crab belonging to the family Retroplumidae.
Archaeopus diabolus[47]	Sp. nov	Valid	Schweitzer et al.	Late Cretaceous		United States
Archaeopus hoploserratus[46]	Sp. nov	Valid	Nyborg et al.	Late Cretaceous (Campanian)	Northumberland Formation	Canada	A crab belonging to the family Retroplumidae.
Archaeopus morenoensis[46]	Sp. nov	Valid	Nyborg et al.	Late Cretaceous (Maastrichtian)	Moreno Formation	United States	A crab belonging to the family Retroplumidae.
Bachmayerus gasparici[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Diogenidae.
Bachmayerus matushyznyi[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Diogenidae.
Bahiacaris[48]	Gen. et comb. nov	Valid	Schweitzer et al.	Cretaceous		Brazil	A caridean shrimp; a new genus for "Atyoida" roxoi Beurlen (1950).
Bathycalliax otsuka[49]	Sp. nov	Valid	Ando, Kawano & Ugai	Eocene	Sakasegawa Formation	Japan	A member of Axiidea belonging to the family Bathycalliacidae.
Bolcacalliax[50]	Gen. et comb. nov	Valid	Pasini et al.	Eocene (Ypresian)		Italy	A member of the family Callianassidae; a new genus for "Protaxius" eocenicus Secrétan (1975).
Brachynotus multituberculatus[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A species of Brachynotus.
Calappilia gableorum[37]	Sp. nov	Valid	Feldmann, Schweitzer & Phillips	Oligocene		United States	A species of Calappilia.
Calappilia granulosa[37]	Sp. nov	Valid	Feldmann, Schweitzer & Phillips	Oligocene		United States	A species of Calappilia.
Calappilia guttata[51]	Sp. nov	Valid	Pasini, Garassino & Damarco	Oligocene (Rupelian)		Italy	A species of Calappilia.
Calappilia perreaulti[37]	Sp. nov	Valid	Feldmann, Schweitzer & Phillips	Oligocene		United States	A species of Calappilia.
Callichimaera[52]	Gen. et sp. nov	Valid	Luque et al.	Late Cretaceous (Cenomanian to Turonian)	Churuvita Group Frontier Formation	Colombia United States	A crab belonging to the new family Callichimaeridae. The type species is C. perplexa.
Caloxanthus kavanaghi[53]	Sp. nov	Valid	Collins, Garvie & Mellish	Cretaceous (Albian/Cenomanian)	Del Rio Formation	United States	A crab belonging to the group Etyoidea and to the family Feldmannidae.
Cenocorystes libbyae[54]	Sp. nov	Valid	Van Bakel et al.	Late Cretaceous (Maastrichtian)	Peedee Formation	United States	A palaeocorystoid crab.
Corallomursia prominens[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Calappidae.
Costadromia[55]	Gen. et sp. nov	Valid	Feldmann & Schweitzer	Late Cretaceous (late Campanian)	Wenonah Formation	United States	A sponge crab. The type species is C. hajzeri.
Cretainermis[56]	Gen. et sp. nov	Valid	Prado, Calado & Barreto	Early Cretaceous (Aptian)	Romualdo Formation	Brazil	A member of Penaeoidea. Genus includes new species C. pernambucensis.
Cretamaja klompmakeri[57]	Sp. nov	Valid	González-León & Moreno-Bedmar in Vega, González-León & Moreno-Bedmar	Early Cretaceous (Barremian)	Zapotitlan Formation	Mexico	A crab.
Cretatrizocheles doerflesensis[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Pylochelidae.
Cymodoce (?) scopula[58]	Sp. nov	Valid	Fraaije et al.	Early Cretaceous (Albian)	Basque-Cantabrian Basin	Spain	A sphaeromatid isopod.
Daragrapsus setalis[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of Grapsoidea of uncertain phylogenetic placement.
Dardanus caporiondoi[59]	Sp. nov	Valid	Ceccon & De Angeli	Early Oligocene		Italy	A species of Dardanus.
Devonoaxius[60]	Gen. et sp. nov	Valid	Feldmann & Schweitzer	Devonian (Famennian)	Ohio Shale	United States	A member of Axiidea. Genus includes new species D. garlandi.
Dinocarcinus[61]	Gen. et sp. nov		Van Bakel, Hyžný, Valentin & Robin in Robin et al.	Late Cretaceous (late Campanian)		France	A crab, a member of Portunoidea sensu lato. The type species is D. velauciensis.
Doraranina[62]	Gen. et sp. nov	Valid	Gustafson, Nyborg & van Bakel	Early Eocene	Lookingglass Formation	United States	A crab belonging to the group Raninoidea and to the family Lyreididae. Genus includes new species D. manleyi.
Dromiopsis marginospinosa[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Dromiidae.
Dromiopsis oscari[63]	Sp. nov	Valid	Nyborg et al.	Early Cretaceous (Albian)	Hudspeth Formation	United States
?Dromiopsis rocchettii[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Dromiidae.
Ebalia pumicosa[59]	Sp. nov	Valid	Ceccon & De Angeli	Early Oligocene		Italy	A species of Ebalia.
Eobodotria[64]	Gen. et sp. nov	Valid	Luque & Gerken	Late Cretaceous (Cenomanian–Turonian)		Colombia	A member of Cumacea. Genus includes new species E. muisca.
Eocarpilius vicetinus[59]	Sp. nov	Valid	Ceccon & De Angeli	Early Oligocene		Italy	A crab belonging to the group Carpilioidea and the family Carpiliidae.
Eodorippe connori[63]	Sp. nov	Valid	Nyborg et al.	Early Cretaceous (Albian)	Hudspeth Formation	United States	A crab belonging to the superfamily Dorippoidea and the family Telamonocarcinidae. Originally described as a species of Eodorippe, but subsequently transferred to the genus Personadorippe.[65]
Eopaguropsis grandis[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Diogenidae.
Eopaguropsis schindewolfi[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Diogenidae.
Eucalliax capsulasetaea[66]	Sp. nov	Valid	Kiel et al.	Early Oligocene	Talara Basin	Peru	A member of the family Callianassidae.
Eumunida veronensis[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A species of Eumunida.
Folguerolesia collinsi[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of Leucosioidea belonging to the family Folguerolesiidae.
?Galathea genesis[67]	Sp. nov	Valid	Robins & Klompmaker	Late Jurassic	Ernstbrunn Limestone	Austria	A member of the family Galatheidae.
Galatheites britmelanarum[67]	Sp. nov	Valid	Robins & Klompmaker	Late Jurassic	Ernstbrunn Limestone	Austria	A member of Galatheoidea belonging to the family Catillogalatheidae.
Gastrosacus raboeufi[68]	Sp. nov	Valid	Fraaije et al.	Middle Jurassic (Bathonian)		France	A member of Galatheoidea.
Glaessnerella yamoori[63]	Sp. nov	Valid	Nyborg et al.	Early Cretaceous (Albian)	Hudspeth Formation	United States
Glabropilumnus bizzarinii[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Pilumnidae.
Glabropilumnus tuberculatus[59]	Sp. nov	Valid	Ceccon & De Angeli	Early Oligocene		Italy	A crab belonging to the group Pilumnoidea and the family Pilumnidae.
Gyroplanus[47]	Gen. et sp. nov	Valid	Schweitzer et al.	Late Cretaceous		United States	A cyclodorippoid crab. Genus includes new species G. sapphirinus.
Hepatiscus meneghinii[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Aethridae.
Hinecaris[38]	Gen. et sp. nov	Valid	Karasawa et al.	Late Cretaceous (Campanian–Maastrichtian)	Izumi Group	Japan	A member of the family Axiidae. The type species is H. simplex.
Kromtitis lluisprietoi[69]	Sp. nov	Valid	Ossó	Eocene (Lutetian)		Spain	A crab belonging to the family Dynomenidae.
Lindahomola[70]	Gen. et sp. nov	Valid	Garassino et al.	Late Cretaceous (Maastrichtian)	Peedee Formation	United States	A crab belonging to the family Homolidae. Genus includes new species L. longispina.
Linuparus shacklefordae[47]	Sp. nov	Valid	Schweitzer et al.	Late Cretaceous		United States	A palinurid lobster.
Lophoranina lincki[37]	Sp. nov	Valid	Feldmann, Schweitzer & Phillips	Oligocene		United States	A crab.
Macroacaena hannya[49]	Sp. nov	Valid	Ando, Kawano & Ugai	Eocene	Sakasegawa Formation	Japan	A crab belonging to the family Lyreididae.
Maja prealpina[59]	Sp. nov	Valid	Ceccon & De Angeli	Early Oligocene		Italy	A species of Maja.
Matutites periosus[37]	Sp. nov	Valid	Feldmann, Schweitzer & Phillips	Eocene		United States	A crab.
Mesoparapylocheles strouhali[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Parapylochelidae.
Mesoparapylocheles zapfei[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Parapylochelidae.
Mesostylus pervesleri[71]	Sp. nov	Valid	Hyžný & Summesberger	Late Cretaceous (Maastrichtian)	Piesting Formation	Austria	A member of the family Callianassidae.
Montemagambrus[72]	Gen. et sp. nov	Valid	Ceccon & De Angeli	Early Eocene		Italy	A crab belonging to the family Parthenopidae. Genus includes new species M. corallinus.
Munidopsis salinaria[73]	Sp. nov	Valid	Gašparič et al.	Miocene (Langhian)	Tuzla Basin	Bosnia and Herzegovina	A species of Munidopsis.
Munitheites kuepperi[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Schobertellidae.
Nachioides[41]	Gen. et sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Inachoididae. The type species is N. tuberculatus.
Natatolana poblana[74]	Sp. nov	Valid	Vega & Bruce in Vega et al.	Early Cretaceous (Valanginian-Hauterivian)	San Juan Raya Formation	Mexico	An isopod belonging to the family Cirolanidae.
Nicolisia[41]	Gen. et sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of Xanthoidea belonging to the family Eumedonidae. The type species is N. antiqua.
Notopus adinae[37]	Sp. nov	Valid	Feldmann, Schweitzer & Phillips	Oligocene		United States	A crab, a species of Notopus.
Ovalopus[67]	Gen. et comb. nov	Valid	Robins & Klompmaker	Late Jurassic	Ernstbrunn Limestone	Austria Czech Republic Romania	A crab belonging to the group Homolodromioidea and to the family Jurellanidae. The type species is O. hoheneggeri (Moericke, 1889); genus also includes "Coelopus" convexus Schweitzer & Feldmann (2010) and "Coelopus" repandus Schweitzer & Feldmann (2010).
Pachycheles bazzanii[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Porcellanidae.
Pagurus haasei[75]	Sp. nov	Valid	Polkowsky & Fraaije	Oligocene (Chattian)		Germany	A hermit crab, a species of Pagurus.
Palaemon aestuarius[76]	Sp. nov		Du et al.	Early Miocene	Mexican amber	Mexico	A species of Palaemon.
Palaeopalinurus serratus[58]	Sp. nov	Valid	Fraaije et al.	Early Cretaceous (Albian)	Basque-Cantabrian Basin	Spain	A spiny lobster.
Palinurus? sauerweinorum[47]	Sp. nov	Valid	Schweitzer et al.	Late Cretaceous		United States	A palinurid lobster.
Panopeus santurbanensis[59]	Sp. nov	Valid	Ceccon & De Angeli	Early Oligocene		Italy	A species of Panopeus.
Paronachoides[41]	Gen. et sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Inachoididae. The type species is P. scaber.
Percnon santurbanensis[77]	Sp. nov	Valid	Ceccon & de Angeli	Oligocene		Italy	A crab, a species of Percnon.
Petrolisthes punctatus[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A species of Petrolisthes.
Pilumnopeus priscus[59]	Sp. nov	Valid	Ceccon & De Angeli	Early Oligocene		Italy	A crab belonging to the group Pilumnoidea and the family Pilumnidae.
Pilumnus pedemontanus[59]	Sp. nov	Valid	Ceccon & De Angeli	Early Oligocene		Italy	A species of Pilumnus.
Planoprosopon ogawaense[78]	Sp. nov	Valid	Karasawa & Hirota	Late Jurassic–Early Cretaceous	Yatsuji Formation	Japan	A crab belonging to the group Homolodromioidea and to the family Longodromitidae.
Planoprosopon sarumaru[78]	Sp. nov	Valid	Karasawa & Hirota	Late Jurassic–Early Cretaceous	Yatsuji Formation	Japan	A crab belonging to the group Homolodromioidea and to the family Longodromitidae.
Pretrizocheles[42]	Gen. et sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Pylochelidae. The type species is P. cocullo.
Prolecythocaris rieberi[79]	Sp. nov	Valid	Schweigert	Late Jurassic		Germany	A crab belonging to the group Dromiacea and to the family Lecythocaridae.
Protopagurus[42]	Gen. et sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Paguridae. The type species is P. janoscheki.
Pseudocarinocarcinoides[41]	Gen. et sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of Goneplacoidea belonging to the family Carinocarcinoididae. The type species is P. karasawai.
Retropluma minuta[73]	Sp. nov	Valid	Gašparič et al.	Miocene (Langhian)	Tuzla Basin	Bosnia and Herzegovina	A species of Retropluma.
Ripleycarcinus[47]	Gen. et sp. nov	Valid	Schweitzer et al.	Late Cretaceous		United States	A homoloid crab. Genus includes new species R. primaevus.
Ripleycorystes[47]	Gen. et sp. nov	Valid	Schweitzer et al.	Late Cretaceous		United States	A palaeocorystoid crab. Genus includes new species R. cardwelli.
Symethoides danieli[47]	Sp. nov	Valid	Schweitzer et al.	Late Cretaceous		United States	A raninoid crab.
Tethysmunida setifera[41]	Sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Munidopsidae.
Tithopaguristes[42]	Gen. et sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Diogenidae. The type species is T. porosus.
Tropidicarcinus[47]	Gen. et 2 sp. nov	Valid	Schweitzer et al.	Late Cretaceous		United States	A dakoticancroid crab. Genus includes new species T. mcanallyae and T. starnesi.
Ululapagurus oroszyi[42]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A hermit crab belonging to the family Diogenidae.
Vacuotheca[80]	Gen. et sp. nov	Valid	Schädel, Perrichot & Haug	Late Cretaceous (Turonian)		France	A member of Isopoda belonging to the group Epicaridea. The type species is V. dupeorum.
Valamunida[81]	Gen. et sp. nov	Valid	Klompmaker & Robins in Hryniewicz et al.	Late Paleocene	Basilika Formation	Norway	A member of the family Munididae. The type species is V. haeggi.
Verarena[41]	Gen. et sp. nov	Valid	Beschin, Busulini & Tessier in Beschin et al.	Eocene (Priabonian)		Italy	A member of the family Euryplacidae. The type species is V. katoi.
Vibrissalana[67]	Gen. et sp. nov	Valid	Robins & Klompmaker	Late Jurassic	Ernstbrunn Limestone	Austria	A porcelain crab. Genus includes new species V. jurassica.
Vicluvia[82]	Gen. et sp. nov	Valid	Larghi et al.	Middle Triassic (Ladinian)	Meride Limestone	Italy Switzerland	A member of Lophogastrida. Genus includes new species V. lombardoae.

Ostracods

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Amicytheridea bilthanaensis[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Bairdiocypris fabiformis[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Bairdoppilata dorsoarcuata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Bairdiidae.
Bairdoppilata pintoi[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	La Rosa Formation Pirabas Formation	Brazil Venezuela	A member of Podocopida belonging to the family Bairdiidae.
Bairdoppilata turonica[85]	Sp. nov	Valid	Slipper	Late Cretaceous (Turonian)		United Kingdom
Bairdoppilata vandenboldi[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Bairdiidae.
Bakunella anae[86]	Sp. nov	Valid	Spadi et al.	Late Miocene–Pliocene		Croatia	A member of the family Candonidae.
Bassleratia torquata[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Bolbozoe psittaca[87]	Sp. nov	Valid	Perrier et al.	Silurian		United Kingdom	A member of Myodocopa.
Boreobolbina morkokiana[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Bradleya majorani[88]	Sp. nov	Valid	Bergue, Brandão & Anjos Zerfass	Pleistocene		Atlantic Ocean (Rio Grande Rise)	A member of the family Thaerocytheridae.
Bradleya ybate[88]	Sp. nov	Valid	Bergue, Brandão & Anjos Zerfass	Late Miocene		Atlantic Ocean (Rio Grande Rise)	A member of the family Thaerocytheridae.
Bythocypris wangi[89]	Sp. nov	Valid	Tanaka, Siveter & Williams	Devonian	Fukuji Formation	Japan
Bythoceratina? asulcata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Bythocytheridae.
Bythoceratina? sinuocostata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Bythocytheridae.
Cardobairdia longitecta[85]	Sp. nov	Valid	Slipper	Late Cretaceous (Turonian)		United Kingdom
Carinobairdia cabralae[90]	Sp. nov		Forel in Forel, Thuy & Wisshak	Late Triassic (Carnian)	Maantang Formation	China	A member of the family Bairdiidae.
Cativella primaveraensis[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Trachyleberididae.
Clavofabellina fukujiensis[89]	Sp. nov	Valid	Tanaka, Siveter & Williams	Devonian	Fukuji Formation	Japan
Clintiella antifrigga[91]	Sp. nov	Valid	Siveter et al.	Silurian	Hitoegane Formation	Japan	A beyrichioid palaeocopid ostracod.
Coelochilina reticulata[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Costa spinaventralis[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Trachyleberididae.
Curfsina coarctata[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Curfsina hanumanpuraensis[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Cushmanidea? bragantinaensis[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cushmanideidae.
Cyprideis? carmoi[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cytherideidae.
Cytherella altacaelateralis[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of the family Cytherellidae.
Cytherella kempfi[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of the family Cytherellidae.
Cytherella notossinuosa[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of the family Cytherellidae.
Cytherella truncatoides[85]	Sp. nov	Valid	Slipper	Late Cretaceous (Turonian)		United Kingdom
Cytherella vulna[85]	Sp. nov	Valid	Slipper	Late Cretaceous (Turonian)		United Kingdom
Cytherella weaveri[85]	Sp. nov	Valid	Slipper	Late Cretaceous (Turonian)		United Kingdom
Cytherelloidea colini[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of the family Cytherellidae.
Cytherelloidea cumpassorobusta[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of the family Cytherellidae.
Cytherelloidea granulosa parca[85]	Subsp. nov	Valid	Slipper	Late Cretaceous (Turonian)		United Kingdom
Cytherelloidea mediocythara[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of the family Cytherellidae.
Cytherelloidea quasilenisa[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of the family Cytherellidae.
Cytheretta fortiscostata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cytherettidae.
Cytheretta petrii[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cytherettidae.
Cytheropteron hanumanpuraensis[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Cytheropteron ratitalaiensis[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Cytherura? ornatareticulum[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cytheruridae.
Cytherura? punctocentrata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cytheruridae.
Cytherura? quasilenisa[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cytheruridae.
Eocytheropteron bilthanaensis[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Gibbosocythere[92]	Gen. et 2 sp. nov	Valid	Karpuk, Whatley & Maybury	Early Cretaceous		Crimean Peninsula	A member of Paradoxostomatidae. Genus includes new species G. ferelevis and G. cellulata.
Hemicytherura bradyisimilis[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cytheruridae.
Hiatobairdia senegasi[90]	Sp. nov		Forel in Forel, Thuy & Wisshak	Late Triassic (Carnian and Norian)	Kuahongdong Formation Maantang Formation	China	A member of the family Bairdiidae.
Hiatobairdia zhengshuyingi[90]	Sp. nov		Forel in Forel, Thuy & Wisshak	Late Triassic (Carnian and Norian)	Kuahongdong Formation Maantang Formation	China	A member of the family Bairdiidae.
Hollinella orienta[91]	Sp. nov	Valid	Siveter et al.	Silurian	Gionyama Formation	Japan	A hollinoid palaeocopid ostracod.
Hulingsina? lorenesmithae[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cushmanideidae.
Hulingsina reticulorugosa[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cushmanideidae.
Hungarella gommerii[90]	Sp. nov		Forel in Forel, Thuy & Wisshak	Late Triassic (Carnian)	Maantang Formation	China	A member of the family Healdiidae.
Jatella ampla[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Krithe araucoensis[93]	Sp. nov	Valid	Bergue, Coimbra & Finger	Early Miocene		Chile
Krithe dawnpetersonae[93]	Sp. nov	Valid	Bergue, Coimbra & Finger	Early Miocene		Chile
Krithe nerudai[93]	Sp. nov	Valid	Bergue, Coimbra & Finger	Early Miocene		Chile
Leperditella paula[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Loxoconcha corpulenta[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Loxoconchidae.
Microcythere acuminata[88]	Sp. nov	Valid	Bergue, Brandão & Anjos Zerfass	Quaternary		Atlantic Ocean (Rio Grande Rise)	A member of the family Microcytheridae.
Moierina grata[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Mydionobairdia cauladelicata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Bairdiidae.
Nigeroloxoconcha baghensis[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Nigeroloxoconcha diluta[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Nudator[87]	Gen. et 4 sp. nov	Valid	Perrier et al.	Silurian (Wenlock and Ludlow)		Poland United Kingdom France?	A member of Myodocopa beloning to the family Entomozoidae. The type species is N. inflatus; genus also includes new species N. angiportatus, N. artumatus and N. elegantulus.[94]
Ochesaarina aculeata[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Oculparva[87]	Gen. et sp. nov	Valid	Perrier et al.	Silurian		United Kingdom	A member of Myodocopa. Genus includes new species O. moncola.
Ovocytheridea baghensis[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Paijenborchella jeerabadensis[83]	Sp. nov	Valid	Chaudhary & Nagori	Late Cretaceous	Bagh Formation	India
Pauproles[91]	Gen. et sp. nov	Valid	Siveter et al.	Silurian	Hitoegane Formation	Japan	A eurychilinoid palaeocopid ostracod. Genus includes new species P. supparata.
Piscarista[87]	Gen. et sp. nov	Valid	Perrier et al.	Silurian		United Kingdom	A member of Myodocopa. Genus includes new species P. sagenata.
Planiglandites[84]	Gen. et sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia	Genus includes new species P. mirabilis'.
Pontocyprella goussardi[90]	Sp. nov		Forel in Forel, Thuy & Wisshak	Late Triassic (Carnian)	Maantang Formation	China	A member of the family Pontocyprididae.
Pontocyprella robusta cometa[85]	Subsp. nov	Valid	Slipper	Late Cretaceous (Turonian)		United Kingdom
Pontocypris aguilerai[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Late Oligocene to early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Pontocypridae.
Pseudotethysia[92]	Gen. et sp. nov	Valid	Karpuk, Whatley & Maybury	Early Cretaceous		Crimean Peninsula	A member of Cytheruridae. Genus includes new species P. reticulata.
Quadracythere adornata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Hemicytheridae.
Quadracythere fortisrobusta[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Hemicytheridae.
Quadracythere limbimodesta[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Hemicytheridae.
Rectella cistelliformis[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Retiprimites alveolatus[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Retiprimites mirandus[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia
Rostrocytheridea decurtata[95]	Sp. nov	Valid	Chaudhary, Nagori & Khosla	Late Cretaceous	Bagh Formation	India	A member of the family Cytherideidae.
Rostrocytheridea divergens[95]	Sp. nov	Valid	Chaudhary, Nagori & Khosla	Late Cretaceous	Bagh Formation	India	A member of the family Cytherideidae.
Semicytherura? fortisreticulata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Cytheruridae.
Seminova[87]	Gen. et comb. nov	Valid	Perrier et al.	Silurian		United Kingdom	A member of Myodocopa belonging to the superfamily Bolbozoidea. The type species is "Entomis" depressa Jones (1884).
Tenedocythere? rugosocostata[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Hemicytheridae.
Touroconcha ishizakii[30]	Sp. nov	Valid	Nogueira, Ramos & Hunt	Early Miocene	Pirabas Formation	Brazil	A member of Podocopida belonging to the family Loxoconchidae.
Uthoernia sibirica[84]	Sp. nov	Valid	Melnikova	Late Ordovician	Delingde Formation	Russia

Other crustaceans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Austromegabalanus carrioli[96]	Sp. nov	Valid	Collareta et al.	Early Miocene	Chilcatay Formation	Peru	A barnacle.
Calmanostraca hassbergella[97]	Sp. nov		Wagner, Hauh & Haug	Late Triassic (Carnian)	Hassberge Formation	Germany	A member of Branchiopoda belonging to the group Calmanostraca.
Cretiscalpellum matrioni[98]	Sp. nov	Valid	Gale	Early Cretaceous (Albian)		France	A barnacle. Originally described as a species of Cretiscalpellum; Gale (2020) transferred it to the genus Witherscalpellum.[99]
Cretiscalpellum mutterlosei[98]	Sp. nov	Valid	Gale	Early Cretaceous (Hauterivian)		Germany	A barnacle.
Halicyne oosterinkorum[100]	Sp. nov	Valid	Schweitzer, Feldmann & Schinker	Middle Triassic (Anisian)		Netherlands	A member of Cyclida.
Jaegerscalpellum[98]	Gen. et sp. et comb. nov	Valid	Gale	Early Cretaceous (Hauterivian to Albian)		Germany	A barnacle. Genus includes new species J. elegans, as well as J. comptum (Withers, 1910) and J. politum (Darwin, 1851).
Martillepas auriculum[98]	Sp. nov	Valid	Gale	Early Cretaceous (Hauterivian)		Germany	A barnacle.
Martillepas decoratus[98]	Sp. nov	Valid	Gale	Early Cretaceous (Hauterivian)		Germany	A barnacle.
Notostraca oleseni[97]	Sp. nov		Wagner, Hauh & Haug	Early Cretaceous	Yixian Formation	China	A member of Branchiopoda belonging to the group Notostraca.
Paraleptestheria mitchelli[101]	Sp. nov	Valid	Gallego et al.	Late Eocene		United Kingdom	A clam shrimp.
Pedupycnolepas pulcher[98]	Sp. nov	Valid	Gale	Early Cretaceous (Hauterivian)		Germany	A barnacle.
Pemphilimnadiopsis cheni[102]	Sp. nov	Valid	Liao et al.	Carboniferous (Pennsylvanian)	Benxi Formation	China	A clam shrimp.
Perumegabalanus[103]	Gen. et sp. nov	Valid	Coletti et al.	Miocene (Burdigalian)	Chilcatay Formation	Peru	A barnacle. Genus includes new species P. calziai.
Platylepas mediterranea[104]	Sp. nov	Valid	Collareta et al.	Early Pleistocene (Gelasian)		Italy	A barnacle, a species of Platylepas.
Prolatcyclus[105]	Gen. et comb. et sp. nov	Valid	Mychko et al.	Carboniferous (Viséan)		Russia United Kingdom	A member of Cyclida. Genus includes P. martinensis (Goldring, 1967) and a new species P. kindzadza.
Verruca alabamensis[106]	Sp. nov	Valid	Perreault & Buckeridge	Paleogene	Yazoo Formation	United States	A barnacle belonging to the family Verrucidae.
Verruca gailgoedertae[106]	Sp. nov	Valid	Perreault & Buckeridge	Middle Eocene		United States	A barnacle belonging to the family Verrucidae.
Verruca sorrellae[106]	Sp. nov	Valid	Perreault & Buckeridge	Late Eocene to earliest Oligocene		United States	A barnacle belonging to the family Verrucidae.

Insects

Trilobites

Research

A study on the rates of evolution of trilobites in the Cambrian is published by Paterson, Edgecombe & Lee (2019), who interpret their findings as indicative of Cambrian rather than Precambrian origin of trilobites, and as indicating that the Cambrian explosion had already largely concluded by the time the typical Cambrian fossil record begins (∼521 Ma).[107]
A study on repaired injuries in trilobite specimens from the Cambrian Ruin Wash Lagerstätte (Nevada, United States) is published by Pates & Bicknell (2019), who interpret their findings as indicative of species specific predation on Cambrian trilobites, and of enlarged spines in some trilobite species serving as possible predation deterrents.[108]
A study on the morphology and ontogeny of Changaspis elongata, based on data from specimens from the Lazizhai section of the Balang Formation (Cambrian Stage 4; Guizhou, China), is published by Du et al. (2019).[109]
A cluster of over a 100 juvenile trilobites belonging to the genus Aphelaspis is reported from the Cambrian (Paibian) Conasauga Formation (Georgia, United States) by Schwimmer & Montante (2019).[110]
A study on patterns of occupancy and diversity trajectories in late Cambrian-early Ordovician trilobite communities from the Argentine Cordillera Oriental is published by Serra, Balseiro & Waisfeld (2019).[111]
A study on the phylogenetic relationships of members of the family Olenidae is published by Monti & Confalonieri (2019).[112]
A study on the phylogenetic relationships of members of the olenid subfamily Balnibarbiinae is published by Hopkins (2019).[113]
Nine specimens of Symphysurus ebbestadi preserving all key stages of the exoskeleton moulting process are described from the Lower Ordovician Fezouata Formation (Morocco) by Drage et al. (2019).[114]
A study on clusters of Ampyx priscus from the Ordovician Fezouata Shale of Morocco is published by Vannier et al. (2019), who interpret these clusters as evidence of a collective behaviour rather than a result of passive transportation and accumulation.[115]
A study on the phylogenetic relationships of the species and subspecies assigned to the phacopid genus Austerops is published by Oudot et al. (2019).[116]
A study on the morphological changes in the ontogeny of the Ordovician phacopid trilobite Calyptaulax strasburgensis is published by Jacobs & Carlucci (2019).[117]
A well-preserved body cluster of 18 specimens of Arctinurus boltoni is described from the Silurian Rochester Shale (New York, United States) by Bicknell, Paterson & Hopkins (2019).[118]
A study on development and evolution of the cephalon and the pygidium in phacopid trilobites is published by Oudot et al. (2019).[119]
A study on the internal structures of eyes of trilobites belonging to the genera Asaphus and Archegonus is published by Scholtz, Staude & Dunlop (2019).[120]
A study on the variability of trilobite moulting behaviour is published by Drage (2019).[121]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Anacaenaspis yanpingensis[122]	Sp. nov	Valid	Wei & Zhan	Silurian (Rhuddanian)	Niuchang Formation	China
Asaphellus intermedius[123]	Sp. nov	Valid	Ghobadi Pour	Ordovician		Iran
Asteropyge boeckae[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium
Asteropyge eonia[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	France
Asteropyge filoxenia[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium
Astycoryphe rugocauda[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Couvin Formation	Belgium	A member of Proetida.
Astycoryphe tureki[125]	Sp. nov	Valid	Mergl & Budil	Devonian (Eifelian)		Czech Republic	A member of Proetida.
Bigotina kelleri[126]	Sp. nov	Valid	Geyer	Cambrian Stage 3	Igoudine Formation	Morocco	A member of Redlichiina belonging to the family Bigotinidae.
Bigotina monningeri[126]	Sp. nov	Valid	Geyer	Cambrian		Morocco	A member of Redlichiina belonging to the family Bigotinidae.
Bigotinops chouberti[126]	Sp. nov	Valid	Geyer	Cambrian		Morocco	A member of Redlichiina belonging to the family Bigotinidae.
Brachymetopus (Conimetopus) alekseevi[127]	Sp. nov	Valid	Mychko in Mychko & Savchuk	Early Permian		Russia
Brevibole minerva[128]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Achegoninae.
Bronteopsis tenuirhachis[129]	Sp. nov	Valid	Ove, Ebbestad & Fortey	Late Ordovician	Povorotnaya Formation	Russia	A member of the family Styginidae.
Carbonoproetus bellona[128]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Bollandiinae.
Carbonoproetus volturnus[128]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Bollandiinae.
Conophrys multituberculatus[123]	Sp. nov	Valid	Ghobadi Pour	Ordovician		Iran
Cyphaspides ammari[130]	Sp. nov	Valid	Chatterton, Gibb & McKellar	Devonian (Eifelian)	Bou Tchrafine Group	Morocco	A member of Proetida belonging to the family Aulacopleuridae.
Cyphaspides nicoleae[130]	Sp. nov	Valid	Chatterton, Gibb & McKellar	Devonian (Eifelian)	Bou Tchrafine Group	Morocco	A member of Proetida belonging to the family Aulacopleuridae.
Cyphaspides pankowskiorum[130]	Sp. nov	Valid	Chatterton, Gibb & McKellar	Devonian (Eifelian)	Bou Tchrafine Group	Morocco	A member of Proetida belonging to the family Aulacopleuridae.
Cyphaspis insolata[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium
Cyphaspis iuxta[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium
Cyphaspis konstantini[131]	Sp. nov	Valid	Alberti	Devonian (Emsian)		Germany	A member of the family Aulacopleuridae.
Cyrtoproetus korni[128]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Achegoninae.
Cyrtoproetus roesenbeckensis[128]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Achegoninae.
Cyrtosymbolinella[132]	Gen. et sp. nov	Valid	Basse & Lemke	Devonian		Germany	A member of the family Phillipsiidae belonging to the subfamily Cyrtosymbolinae. The type species is C. svojtkai.
Debrenella[126]	Gen. et sp. nov	Valid	Geyer	Cambrian		Morocco	A member of Redlichiida belonging to the family Fallotaspididae. Genus includes new species D. larvalis.
Diademaproetus pertinax[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium	A member of Proetida.
Dionide trigintasegmentata[129]	Sp. nov	Valid	Ove, Ebbestad & Fortey	Late Ordovician	Barkov Formation	Russia	A member of Asaphida belonging to the family Dionididae.
Eladiolinania[126]	Gen. et 2 sp. et comb. nov	Valid	Geyer	Cambrian	Igoudine Formation	Morocco Spain?	A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species E. castor and E. pollux, and possibly also "Serrania" palaciosi Liñán et al. (2008) and "Serrania" gordaensis Liñán et al. (2008).
Evenkaspis (Bilobaspis) mirabilis[129]	Subgen. et sp. nov	Valid	Ove, Ebbestad & Fortey	Late Ordovician	Tolmachev Formation	Russia	A member of the family Pterygometopidae.
Failleana superba[129]	Sp. nov	Valid	Ove, Ebbestad & Fortey	Late Ordovician	Povorotnaya Formation	Russia	A member of the family Styginidae.
Fallotaspis antecedens[126]	Sp. nov	Valid	Geyer	Cambrian	Igoudine Formation	Morocco	A member of Redlichiida belonging to the family Fallotaspididae.
Geesops icovellaunae[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium
Gerastos kesselaeri[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium
Gerastos silvicultrix[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Emsian)	Heisdorf Formation	Germany
Heliopeltis morrisoni[133]	Sp. nov	Valid	Basse, Müller & Habenicht	Devonian (Emsian)		Germany	A member of the family Scutelluidae belonging to the subfamily Kolihapeltinae.
Hollandiella[134]	Gen. et 3 sp. nov	Valid	Van Viersen & Lerouge	Devonian (Pragian and Emsian)	Ihandar Formation Timrhanrhart Formation	Morocco	A member of the family Proetidae belonging to the subfamily Cornuproetinae. The type species is H. curvirostris; genus also includes new species H. lebruni and H. verecunda.
Interproetus mizobuchii[135]	Sp. nov	Valid	Stocker et al.	Paleozoic		Japan	A member of Proetida belonging to the family Proetidae.
Issendalenia[126]	Gen. et sp. nov	Valid	Geyer	Cambrian	Igoudine Formation	Morocco	A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species I. grandispina.
Kasachstania alperovichi[136]	Sp. nov	Valid	Ghobadi Pour et al.	Devonian (Lochkovian)	Shakshagaily Formation	Kazakhstan
Kellerwaldops[132]	Gen. et sp. nov	Valid	Basse & Lemke	Devonian		Germany	A member of the family Phacopidae belonging to the subfamily Phacopinae. The type species is K. kaufmanni.
Kettneraspis eftychia[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	France	A member of the family Odontopleuridae.
Koneprusia chimaera[131]	Sp. nov	Valid	Alberti	Devonian (Emsian)		Germany	A member of the family Odontopleuridae.
Koneprusites oresibios[134]	Sp. nov	Valid	Van Viersen & Lerouge	Devonian (Eifelian)		Morocco	A member of the family Proetidae belonging to the subfamily Cornuproetinae.
Lepidoproetus arenicolus[134]	Sp. nov	Valid	Van Viersen & Lerouge	Devonian (Pragian)		Morocco	A member of the family Proetidae belonging to the subfamily Cornuproetinae.
Liobole (Liobole) aurora[128]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Achegoninae.
Liobole (Panibole) neptun[128]	Sp. nov	Valid	Müller & Hahn	Carboniferous (Mississippian)		Germany	A member of the family Phillipsiidae belonging to the subfamily Achegoninae.
Nephranops (Maternia)[137]	Subgen. et sp. nov	Valid	Feist	Devonian (Famennian)		France Germany	A member of Phacopida. The type species of the subgenus is "Nephranops incisus" dillanus Richter & Richter (1926) (raised to the rank of a separate species Nephranops (Maternia) dillanus); the subgenus also includes Nephranops franconicus Alberti (1970) and a new species Nephranops (Maternia) occitanicus.
Nephranops (Nephranops) angerae[137]	Sp. nov	Valid	Feist	Devonian (Famennian)		France	A member of Phacopida.
Niobe (Niobella) obscura[138]	Sp. nov	Valid	Zhou & Zhou	Ordovician	Dawan Formation	China
Paralejurus colossicus[139]	Sp. nov	Valid	Alberti	Devonian (Emsian)		Germany	A member of the family Scutelluidae.
Paralejurus flickorum[139]	Sp. nov	Valid	Alberti	Devonian (Emsian)		Germany	A member of the family Scutelluidae.
Pararemopleurides ornatissimus[129]	Sp. nov	Valid	Ove, Ebbestad & Fortey	Late Ordovician	Povorotnaya Formation	Russia	A member of Asaphida belonging to the family Remopleurididae.
Pedinopariops ceuthonymus[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	France	A member of the family Phacopidae.
Pedinopariops richterianus geminus[124]	Subsp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium	A member of the family Phacopidae.
Placoparia africana[140]	Sp. nov	In press	Pereira & Gutiérrez-Marco in Gutiérrez-Marco et al.	Ordovician (Sandbian)		Morocco	Announced in 2019; the final version of the article naming it is not published yet.
Polypleuraspidella[141]	Gen. et comb. nov	Valid	Peel	Cambrian	Spence Shale	United States ( Utah)	A new genus for "Polypleuraspis" cooperi Robison & Babcock (2011).
Polypleuraspis glacialis[141]	Sp. nov	Valid	Peel	Cambrian (Miaolingian)	Telt Bugt Formation	Greenland
Pseudobigotina[126]	Gen. et sp. nov	Valid	Geyer	Cambrian		Morocco	A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species P. antiatlasensis.
Pulvinocephalus[137]	Gen. et comb. et 2 sp. et subsp. nov	Valid	Feist	Devonian (Famennian)		Germany Russia	A member of Phacopida. The type species is "Phacops (Trimerocephalus)" steinachensis Richter & Richter (1926) (including a new subspecies Pulvinocephalus steinachensis triangulus); genus also includes "Phacops (Cryphops)" latilimbatus Maximova (1955) and new species Pulvinocephalus ovatus and Pulvinocephalus walliseri.
Radiaspis taghonorum[142]	Sp. nov	Valid	Van Viersen & Magrean	Middle Devonian	Hanonet Formation	Belgium	A member of the family Odontopleuridae.
Raymondella plastron[129]	Sp. nov	Valid	Ove, Ebbestad & Fortey	Late Ordovician	Povorotnaya Formation	Russia	A member of the family Raphiophoridae.
Redlichia rex[143]	Sp. nov	Valid	Holmes, Paterson & García-Bellido	Cambrian Stage 4	Emu Bay Shale	Australia
Robergia nikolaiseni[129]	Sp. nov	Valid	Ove, Ebbestad & Fortey	Late Ordovician	Barkov Formation	Russia	A member of Asaphida belonging to the family Remopleurididae.
Robergia subtilis[129]	Sp. nov	Valid	Ove, Ebbestad & Fortey	Late Ordovician	Povorotnaya Formation	Russia	A member of Asaphida belonging to the family Remopleurididae.
Scabriscutellum archinalae[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium	A member of the family Scutelluidae.
Sculptoproetus raki[134]	Sp. nov	Valid	Van Viersen & Lerouge	Devonian (Emsian)		Morocco	A member of the family Proetidae belonging to the subfamily Cornuproetinae.
Septimopeltis akatastasia[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium	A member of the family Scutelluidae.
Struveops wiedaensis[137]	Sp. nov	Valid	Feist	Devonian (Famennian)		Germany	A member of Phacopida.
Suvorovaella[126]	Gen. et comb. nov	Valid	Geyer	Cambrian	El'gyan Formation	Russia	A member of Redlichiina belonging to the family Bigotinidae. The type species is "Bigotinops" privus Suvorova (1960); genus might also include "Bigotinops" patrius Suvorova (1960).
Symphysurus ebbestadi[144]	Sp. nov	Valid	Gutierrez-Marco, Rabano & Garcia-Bellido	Ordovician (Tremadocian)		Morocco	A member of the family Nileidae.
Tafilaltaspis rufa[134]	Sp. nov	Valid	Van Viersen & Lerouge	Devonian (Eifelian)		Morocco	A member of the family Proetidae belonging to the subfamily Eremiproetinae.
Tesselacauda kriegerae[145]	Sp. nov	Valid	Adrain & Karim	Early Ordovician	Fillmore Formation Garden City Formation	United States
Tesselacauda morrisoni[145]	Sp. nov	Valid	Adrain & Karim	Early Ordovician	Fillmore Formation	United States
Timsaloproetus gibbae[134]	Sp. nov	Valid	Van Viersen & Lerouge	Devonian (Emsian)	Khebchia Formation	Morocco	A member of the family Proetidae belonging to the subfamily Cornuproetinae.
Timsaloproetus pulchistriatus[134]	Sp. nov	Valid	Van Viersen & Lerouge	Devonian (Emsian)	Timrhanrhart Formation	Morocco	A member of the family Proetidae belonging to the subfamily Cornuproetinae.
Timsaloproetus weddigei[134]	Sp. nov	Valid	Van Viersen & Lerouge	Devonian (Eifelian)	Bicken-Ense Formation	Germany	A member of the family Proetidae belonging to the subfamily Cornuproetinae.
Tioutella[126]	Gen. et sp. nov	Valid	Geyer	Cambrian	Igoudine Formation	Morocco	A member of Redlichiina belonging to the family Bigotinidae. Genus includes new species T. floccofavosa.
Trifoliops septimanicus[137]	Sp. nov	Valid	Feist	Devonian (Famennian)		France	A member of Phacopida.
Tropidocoryphe insciens[124]	Sp. nov	Valid	Van Viersen, Taghon & Magrean	Devonian (Eifelian)	Jemelle Formation	Belgium	A member of Proetida.

Other arthropods

Research

A study on the morphology of appendages of the euarthropod Ercaicunia multinodosa from the Early Cambrian Chengjiang biota (China) is published by Zhai et al. (2019).[146]
Description of the appendicular morphology of Sinoburius lunaris is published by Chen et al. (2019).[147]
Description of a new specimen of Emeraldella brutoni from the Cambrian Wheeler Formation (Drum Mountains, Utah, United States), providing new information on the appendicular anatomy of this species, is published by Lerosey-Aubril & Ortega-Hernández (2019).[148]
Description of new bradoriid fossils from the Chengjiang biota, revealing hitherto unknown morphological differences in limb anatomy between bradoriid species, is published by Zhai et al. (2019), who evaluate the implications of this finding for the knowledge of the phylogenetic relationships of bradoriids.[149]
A study on the appendage structure of Naraoia spinosa at both juvenile and adult stages is published by Zhai et al. (2019).[150]
Description of exceptionally preserved soft tissues from mature individuals of Peronopsis and Ptychagnostus from the Cambrian Burgess Shale (British Columbia, Canada), and a study on their implications for inferring the phylogenetic placement of agnostinids within Arthropoda, is published by Moysiuk & Caron (2019).[151]
Completely enrolled, phosphatized specimens of Tsunyidiscus yanjiazhaiensis are described from Cambrian Stage 3 of South China by Dai et al. (2019).[152]
The ontogenetic series of Pagetia vinusta is documented from the Cambrian (Wuliuan) Kaili Formation (Guizhou, China) by Cui et al. (2019).[153]
A study reevaluating crystals of a copper sulfide mineral found in association with specimens of Marrella splendens from the Burgess Shale, originally interpreted as evidence that the animal's blood contained the Cu-bearing protein hemocyanin, is published by Gaines et al. (2019).[154]
A study on the anatomy of the nervous system of Alalcomenaeus is published by Ortega-Hernández, Lerosey-Aubril & Pates (2019).[155]
Juvenile specimens of stylonurine eurypterids belonging to the family Hardieopteridae are described from the Famennian locality of Strud (Belgium) by Lamsdell et al. (2019), who interpret their findings as evidence indicating that eurypterids underwent a marine to freshwater transition during the Devonian, and that juvenile eurypterids inhabited sheltered nursery pools (and migrated to rivers upon reaching maturity).[156]
A study on the structure of the eyes of eurypterids is published by Schoenemann, Poschmann & Clarkson (2019).[157]
A study on ontogenetic changes in prosomal morphology in Carboniferous horseshoe crab Euproops danae is published by Tashman, Feldmann & Schweitzer (2019).[158]
A study on the anatomy of the appendages of the Cretaceous xiphosurid Tachypleus syriacus and on the evolution of fossil horseshoe crab appendages is published by Bicknell et al. (2019).[159]
A study re-evaluating the fossil evidence for lateral compound eyes in Paleozoic horseshoe crabs is published by Bicknell, Amati & Ortega-Hernández (2019).[160]
A study on the morphology of the millipede Phryssonotus burmiticus from the Cretaceous amber from Myanmar is published by Su, Cai & Huang (2019).[161]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Albalimulus[162]	Gen. et sp. nov		Bicknell & Pates	Carboniferous (Tournaisian)	Ballagan Formation	United Kingdom	A member of Limuloidea. The type species is A. bottoni.
Andrognathus burmiticus[163]	Sp. nov	Valid	Moritz & Wesener	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A millipede, a species of Andrognathus.
Ankitokazocaris tatensis[164]	Sp. nov	Valid	Ehiro et al.	Early Triassic (Olenekian)	Osawa Formation	Japan	A thylacocephalan.
Beckwithia? gracilis[165]	Sp. nov	Valid	Zhu, Lerosey-Aubril & Ortega-Hernández	Cambrian (Furongian)	Sandu Formation	China	A member of Aglaspidida.
Burmanopetalum[166]	Gen. et sp. nov	Valid	Stoev, Moritz & Wesener	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A millipede belonging to the group Callipodida and to the new family Burmanopetalidae. The type species is B. inexpectatum.
Cambroraster falcatus[167]	Gen. et sp. nov	Valid	Moysiuk & Caron	Cambrian (Wuliuan)	Burgess Shale	Canada	A member of Radiodonta belonging to the family Hurdiidae.
Concavicaris parva[164]	Sp. nov	Valid	Ehiro et al.	Early Triassic (Olenekian)	Osawa Formation	Japan	A thylacocephalan.
Duraznovis[168]	Gen. et sp. nov	Valid	Lara et al.	Late Triassic (Carnian)	Potrerillos Formation	Argentina	An arthropod of uncertain phylogenetic placement. Genus includes new species D. gallegoi. It was argued to be an insect belonging to the group Cicadomorpha by Fu & Huang (2020).[169]
Electrosminthuridia[170]	Gen. et sp. nov		Robin, D’Haese & Barden	Miocene	Dominican amber	Dominican Republic	A springtail belonging to the family Sminthurididae. The type species is E. helibionta.
Falcatacaris[171]	Gen. et sp. nov	Valid	Braig et al.	Late Jurassic (Tithonian)	Altmühltal Formation	Germany	A thylacocephalan. The type species is F. bastelbergeri.
Fibulacaris[172]	Gen. et sp. nov	Valid	Izquierdo-López & Caron	Cambrian (Wuliuan)	Burgess Shale	Canada	A bivalved arthropod belonging to the group Mandibulata. The type species is F. nereidis.
Hoplitaspis[173]	Gen. et sp. nov	Valid	Lamsdell, Gunderson & Meyer	Late Ordovician	Big Hill Formation	United States	A member of Chasmataspidida belonging to the family Diploaspididae. The type species is H. hiawathai.
Hyleoglomeris groehni[174]	Sp. nov	Valid	Wesener	Eocene	Baltic amber	Europe (Baltic Sea region)	A millipede belonging to the family Glomeridae.
Ligulacaris[175]	Gen. et sp. nov	Valid	Charbonnier et al.	Early Triassic		United States	A thylacocephalan. Genus includes new species L. parisiana.
Luohuilinella deletres[176]	Sp. nov	Valid	Hou et al.	Cambrian Stage 3	Chiungchussu Formation	China	A probable member of Artiopoda and the order Xandarellida.
Micragnostus iturbensis[177]	Sp. nov	Valid	Tortello & Esteban	Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A member of Agnostida. Announced in 2019; the final version of the article naming it was published in 2020.
Miyagicaris[164]	Gen. et sp. nov	Valid	Ehiro et al.	Early Triassic (Olenekian)	Osawa Formation	Japan	A thylacocephalan. Genus includes new species M. costata.
Mollisonia plenovenatrix[178]	Sp. nov	Valid	Aria & Caron	Cambrian (Wuliuan)	Burgess Shale	Canada	A member of Chelicerata belonging to the family Mollisoniidae.
Mongolarachne chaoyangensis[179]	Sp. nov	Disputed	Cheng et al.	Early Cretaceous	Yixian Formation	China	An arthropod of uncertain phylogenetic placement. Originally described as a spider; Selden et al. (2019) reinterpreted it as a faked fossil based on a fossil crayfish, and considered it to be a probable junior synonym of Cricoidoscelosus aethus.[180]
Parisicaris[175]	Gen. et sp. nov	Valid	Charbonnier et al.	Early Triassic		United States	A thylacocephalan. Genus includes new species P. triassica.
Pickettia[181]	Gen. et comb. nov	Valid	Bicknell, Lustri & Brougham	Late Devonian		United States	A member of Xiphosura; a new genus for "Bellinurus" carteri Eller, 1940.
Pseudobeyrichona monile[182]	Sp. nov	Valid	Streng & Geyer	Cambrian	Tannenknock Formation	Germany	A member of Bradoriida belonging to the family Hipponicharionidae.
Siphonophora hui[183]	Sp. nov	Valid	Jiang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A millipede belonging to the group Siphonophorida.
Sloveniolimulus[184]	Gen. et sp. nov	Valid	Bicknell et al.	Middle Triassic (Anisian)	Strelovec Formation	Slovenia	A horseshoe crab. Genus includes new species S. rudkini.
Stilpnocephalus[185]	Gen. et sp. nov	Valid	Selden, Simonetto & Marsiglio	Carboniferous (Kasimovian)	Meledis Formation	Italy	A horseshoe crab belonging to the family Belinuridae. The type species is S. pontebbanus.
Tasmaniolimulus[186]	Gen. et sp. nov	Valid	Bicknell	Late Permian	Jackey Shale	Australia	A member of Xiphosurida belonging to the family Austrolimulidae. The type species is T. patersoni.
Tuzoia lazizhaiensis[187]	Sp. nov	Valid	Wen et al.	Cambrian Stage 4	Balang Formation	China
Zhiwenia[188]	Gen. et sp. nov	Valid	Du et al.	Cambrian Stage 3	Xiaoshiba Lagerstätte	China	A non‐trilobite artiopodan. Genus includes new species Z. coronata.

References

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[2] Johan Lindgren; Dan-Eric Nilsson; Peter Sjövall; Martin Jarenmark; Shosuke Ito; Kazumasa Wakamatsu; Benjamin P. Kear; Bo Pagh Schultz; René Lyng Sylvestersen; Henrik Madsen; James R. LaFountain Jr; Carl Alwmark; Mats E. Eriksson; Stephen A. Hall; Paula Lindgren; Irene Rodríguez-Meizoso; Per Ahlberg (2019). "Fossil insect eyes shed light on trilobite optics and the arthropod pigment screen". Nature. 573 (7772): 122–125. Bibcode:2019Natur.573..122L. doi:10.1038/s41586-019-1473-z. PMID 31413368. S2CID 199577084.

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[4] Richard J. Howard; Gregory D. Edgecombe; David A. Legg; Davide Pisani; Jesus Lozano-Fernandez (2019). "Exploring the evolution and terrestrialization of scorpions (Arachnida: Scorpiones) with rocks and clocks" (PDF). Organisms Diversity & Evolution. 19 (1): 71–86. doi:10.1007/s13127-019-00390-7.

[5] Raman Patel; Rajendra Singh Rana; Paul A. Selden (2019). "An orb-weaver spider (Araneae, Araneidae) from the early Eocene of India" (PDF). Journal of Paleontology. 93 (1): 98–104. doi:10.1017/jpa.2018.71. S2CID 115144220. Archived from the original (PDF) on 2019-04-29.

[6] Ivan L. F. Magalhaes; Guilherme H. F. Azevedo; Peter Michalik; Martín J. Ramírez (2019). "The fossil record of spiders revisited: implications for calibrating trees and evidence for a major faunal turnover since the Mesozoic". Biological Reviews. 95 (1): 184–217. doi:10.1111/brv.12559. PMID 31713947.

[Caestaranea-7] Paul A. Selden; Diying Huang; Russell J. Garwood (2019). "New spiders (Araneae: Palpimanoidea) from the Jurassic Yanliao Biota of China". Journal of Systematic Palaeontology. 18 (2): 137–185. doi:10.1080/14772019.2019.1584831. S2CID 91199691.

[JSPSabrouxetal-8] Romain Sabroux; Denis Audo; Sylvain Charbonnier; Laure Corbari; Alexandre Hassanin (2019). "150-million-year-old sea spiders (Pycnogonida: Pantopoda) of Solnhofen" (PDF). Journal of Systematic Palaeontology. 17 (22): 1927–1938. doi:10.1080/14772019.2019.1571534. S2CID 91265879.

[9] Lorena Palencia; Enrique Peñalver; Carlos E. Prieto; Francisco José Poyato-Ariza (2019). "First fossil harvestmen (Arachnida: Opiliones) from Spain and notes on the fossil record of Opiliones" (PDF). Palaeontologia Electronica. 22 (1): Article number 22.1.5. doi:10.26879/855. Archived from the original (PDF) on 2019-04-29.

[10] Ekaterina A. Sidorchuk; Marta Konikiewicz; W. Calvin Welbourn; Joanna Mąkol (2019). "Active postlarval forms of plume-footed Eatoniana (Trombidiformes: Parasitengona, Erythraeidae) in the Eocene Baltic amber". Zootaxa. 4647 (1): 44–53. doi:10.11646/zootaxa.4647.1.6. PMID 31716974.

[BzA12-11] Jörg Wunderlich (2019). "What is a spider? Cretaceous fossils modify strongly phylogenetics as well as diagnoses of families, superfamilies and even suborders of spiders (Araneida) and other arthropods" (PDF). In Jörg Wunderlich (ed.). Beiträge zur Araneologie, 12. pp. 1–32.

[Paragilarovia-12] Mohamed Elsaka; Plamen G. Mitov; Jason A. Dunlop (2019). "New fossil harvestmen (Arachnida: Opiliones) in the HOFFEINS amber collection". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 292 (2): 155–169. doi:10.1127/njgpa/2019/0815.

[Jinjumegops-13] Tae-Yoon S. Park; Kye-Soo Nam; Paul A. Selden (2019). "A diverse new spider (Araneae) fauna from the Jinju Formation, Cretaceous (Albian) of Korea" (PDF). Journal of Systematic Palaeontology. 17 (15): 1271–1297. doi:10.1080/14772019.2018.1525441. S2CID 92137182. Archived from the original (PDF) on 2019-04-29.

[14] Paul A. Selden; Torsten Wappler (2019). "A new spider (Arachnida: Araneae) from the Middle Eocene Messel Maar, Germany". Palaeoentomology. 2 (6): 596–601. doi:10.11646/palaeoentomology.2.6.10.

[15] Miguel Ángel García-Villafuerte (2019). "Metagonia esquincacanoi sp. nov.: el primer registro fósil de la familia Pholcidae (Arachnida: Araneae) en el ámbar de Chiapas, México". Revista Mexicana de Biodiversidad. 90 (1): e902927. doi:10.22201/ib.20078706e.2019.90.2927.

[16] Antonio Arillo; Luis S. Subías; Gino Chaves da Rocha; Dany Azar (2019). "First fossil oribatid mite from Lebanese amber (Acariformes, Oribatida, Neoliodidae)". Palaeoentomology. 2 (6): 611–617. doi:10.11646/palaeoentomology.2.6.12.

[17] Andrés O. Porta; Daniel N. Proud; Ezequiel Franchi; Willians Porto; María Bernarda Epele; Peter Michalik (2019). "The first record of caeculid mites from the Cretaceous amber of Myanmar with notes on the phylogeny of the family". Zootaxa. 4647 (1): 23–43. doi:10.11646/zootaxa.4647.1.5. PMID 31716973.

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[19] Adiël A. Klompmaker; Matúš Hyžný; Roger W. Portell; Clément Jauvion; Sylvain Charbonnier; Shane S. Fussell; Aaron T. Klier; Raymond Tejera; Sten Jakobsen (2019). "Muscles and muscle scars in fossil malacostracan crustaceans". Earth-Science Reviews. 194: 306–326. Bibcode:2019ESRv..194..306K. doi:10.1016/j.earscirev.2019.04.012.

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[22] Olga Alcântara Barros; João Hermínio Silva; Gilberto Dantas Saraiva; Bartolomeu Cruz Viana; Alexandre Rocha Paschoal; Paulo Tarso Cavalcante Freire; Naiara Cipriano Oliveira; Amauri Jardim Paula; Maria Somália Viana (2019). "Physicochemical investigation of shrimp fossils from the Romualdo and Ipubi formations (Araripe Basin)". PeerJ. 7: e6323. doi:10.7717/peerj.6323. PMC 6378090. PMID 30783565.

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[25] Joachim T. Haug; Carolin Haug; Günter Schweigert (2019). "The oldest "intermetamorphic" larva of an achelatan lobster from the Lower Jurassic Posidonia Shale, South Germany". Acta Palaeontologica Polonica. 64 (4): 685–692. doi:10.4202/app.00627.2019.

[26] Matúš Hyžný; Giovanni Pasini; Alessandro Garassino (2019). "Supergiants in Europe: on the cirolanid isopod Bathynomus A. MILNE EDWARDS, 1879 (Malacostraca, Peracarida) from the Plio-Pleistocene of Italy". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 291 (3): 283–298. doi:10.1127/njgpa/2019/0802.

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[28] Junjun Song; Cheng Huang; Yiming Gong (2019). "Response of Ostracods (Crustacea) to the Devonian F—F event: Evidence from the Yangdi and Nandong sections in Guangxi, South China". Global and Planetary Change. 173: 109–120. Bibcode:2019GPC...173..109S. doi:10.1016/j.gloplacha.2018.12.015.

[29] Xincheng Qiu; Li Tian; Kui Wu; Michael J. Benton; Dongying Sun; Hao Yang; Jinnan Tong (2019). "Diverse earliest Triassic ostracod fauna of the non‐microbialite‐bearing shallow marine carbonates of the Yangou section, South China" (PDF). Lethaia. 52 (4): 583–596. doi:10.1111/let.12332.

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[35] Larry D. Taylor; Aaron O’Dea; Timothy J. Bralower; Seth Finnegan (2019). "Isotopes from fossil coronulid barnacle shells record evidence of migration in multiple Pleistocene whale populations". Proceedings of the National Academy of Sciences of the United States of America. 116 (15): 7377–7381. doi:10.1073/pnas.1808759116. PMC 6462050. PMID 30910962.

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[39] Antonio de Angeli; Fabio Caporiondo (2019). "Alontecarcinus buratoi n. gen., n. sp. (Decapoda, Brachyura, Potamonidae) un nuovo crostaceo d'acqua dolce dell'Eocene (Bartoniano) di Alonte (Monti Berici, Vicenza, Italia settentrionale)" (PDF). Bollettino del Museo Civico di Storia Naturale di Verona. 43: 5–16.

[40] Hiroaki Karasawa; Masaaki Ohara (2019). "Establishment of a new genus for Callianassa (s.l.) sakakuraorum Karasawa, 2000 (Decapoda: Axiidea)" (PDF). Bulletin of the Mizunami Fossil Museum. 45: 33–42. Archived from the original (PDF) on 2019-04-29.

[Nachioides-41] Claudio Beschin; Alessandra Busulini; Giuliano Tessier; Roberto Zorzin (2019). "La fauna di crostacei dell'Eocene superiore di Parona di Verona (Italia nordorientale): nuovi ritrovamenti". Bollettino del Museo di Storia Naturale di Venezia. 70: 71–142.

[Pretrizocheles-42] René H. B. Fraaije; Cristina Robins; Barry W. M. van Bakel; John W. M. Jagt; Friedrich Bachmayer (2019). "Paguroid anomurans from the Tithonian Ernstbrunn Limestone, Austria – the most diverse extinct paguroid assemblage on record" (PDF). Annalen des Naturhistorischen Museums in Wien, Serie A. 121: 257–289. JSTOR 26595692. Archived from the original (PDF) on 2019-01-17.

[43] Carrie E. Schweitzer; Rodney M. Feldmann; Hiroaki Karasawa; Javier Luque; Francisco J. Vega (2019). "New decapod crustacean (Brachyura, Raninoida) from the Lower Cretaceous Hudspeth Formation of Oregon, USA" (PDF). Bulletin of the Mizunami Fossil Museum. 45: 15–22. Archived from the original (PDF) on 2019-04-29.

[44] Fernando A. Ferratges; Samuel Zamora; Marcos Aurell (2019). "A new genus and species of Parthenopidae MacLeay, 1838 (Decapoda: Brachyura) from the lower Eocene of Spain". Journal of Crustacean Biology. 39 (3): 303–311. doi:10.1093/jcbiol/ruz014.

[45] Danièle Guinot; Gerardo Carbot-Chanona; Francisco J. Vega (2019). "Archaeochiapasidae n. fam., a new early Cenomanian brachyuran family from Chiapas, Mexico, new hypothesis on Lecythocaridae Schweitzer & Feldmann, 2009, and phylogenetic implications (Crustacea, Decapoda, Brachyura, Eubrachyura)" (PDF). Geodiversitas. 41 (7): 285–322. doi:10.5252/geodiversitas2019v41a7. S2CID 146631940. Archived from the original (PDF) on 2019-03-28.

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[NJGPA2931-49] Yusuke Ando; Shigenori Kawano; Hiroaki Ugai (2019). "Two new species of Decapoda from the Eocene Sakasegawa Formation in Amakusa Islands, Kyushu, Japan". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 293 (1): 57–66. doi:10.1127/njgpa/2019/0830.

[50] Giovanni Pasini; Alessandro Garassino; Antonio De Angeli; Matúš Hyžný; Luca Giusberti; Roberto Zorzin (2019). "Eocene decapod faunas from the Konservat-Lagerstätten laminites of "Pesciara" (Bolca, Verona) and Monte Postale (Altissimo, Vicenza) in northeast Italy: a review and update". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 293 (3): 233–270. doi:10.1127/njgpa/2019/0840.

[51] Giovanni Pasini; Alessandro Garassino; Piero Damarco (2019). "A new calappid species from the Ligure Piemontese Basin (NW Italy) and reappraisal of the fossil calappid specimens in the Museo "G. Maini", Ovada". Boletín de la Sociedad Geológica Mexicana. 71 (1): 219–227. doi:10.18268/BSGM2019v71n1a12.

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[CRisopodFraaijeetal-58] René H.B. Fraaije; Mikel A. López-Horgue; Niel L. Bruce; Barry W.M. van Bakel; Pedro Artal; John W.M. Jagt; Adiël A. Klompmaker (2019). "New isopod and achelatan crustaceans from mid–Cretaceous reefal limestones in the Basque-Cantabrian Basin, northern Spain". Cretaceous Research. 101: 61–69. doi:10.1016/j.cretres.2019.04.012.

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[61] Ninon Robin; Barry W. M. van Bakel; Matúš Hyžný; Aude Cincotta; Géraldine Garcia; Sylvain Charbonnier; Pascal Godefroit; Xavier Valentin (2019). "The oldest freshwater crabs: claws on dinosaur bones". Scientific Reports. 9 (1): Article number 20220. Bibcode:2019NatSR...920220R. doi:10.1038/s41598-019-56180-w. PMC 6934782. PMID 31882600.

[62] Eric P. Gustafson; Torrey Nyborg; Barry W.M. van Bakel (2019). "A new genus and species of raninid crab, Doraranina manleyi (Decapoda: Raninoidea: Lyreididae) from the lower Eocene of Oregon". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 292 (2): 121–128. doi:10.1127/njgpa/2019/0811.

[NJGPA2932Nyborgetal-63] Torrey Nyborg; Alessandro Garassino; Francisco J. Vega; Gregory Kovalchuk (2019). "Fossil decapod crustaceans from the Early Cretaceous (Albian) of Oregon, USA". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 293 (2): 127–138. doi:10.1127/njgpa/2019/0836.

[64] Javier Luque; Sarah Gerken (2019). "Exceptional preservation of comma shrimp from a mid-Cretaceous Lagerstätte of Colombia, and the origins of crown Cumacea". Proceedings of the Royal Society B: Biological Sciences. 286 (1916): Article ID 20191863. doi:10.1098/rspb.2019.1863. PMC 6939266. PMID 31771467.

[65] Barry W.M. van Bakel; Eduard V. Mychko; Andrej Spiridonov; John W.M. Jagt; René H.B. Fraaije (2020). "New Cretaceous crabs (Crustacea, Brachyura) from Moscow Oblast and Dagestan (Russia): patterns in phylogeny and morphospace of the oldest eubrachyurans (Dorippoidea)". Cretaceous Research. 119: Article 104675. doi:10.1016/j.cretres.2020.104675.

[66] Steffen Kiel; Frida Hybertsen; Matúš Hyžný; Adiël A. Klompmaker (2020). "Mollusks and a crustacean from early Oligocene methane-seep deposits in the Talara Basin, northern Peru". Acta Palaeontologica Polonica. 65 (1): 109–138. doi:10.4202/app.00631.2019.

[Vibrissalana-67] Cristina M. Robins; Adiël A. Klompmaker (2019). "Extreme diversity and parasitism of Late Jurassic squat lobsters (Decapoda: Galatheoidea) and the oldest records of porcellanids and galatheids". Zoological Journal of the Linnean Society. 187 (4): 1131–1154. doi:10.1093/zoolinnean/zlz067.

[68] René H.B. Fraaije; Barry W.M. Van Bakel; John W.M. Jagt; Sylvain Charbonnier; Jean-Philippe Pezy (2019). "The oldest record of galatheoid anomurans (Decapoda, Crustacea) from Normandy, northwest France". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 292 (3): 291–297. doi:10.1127/njgpa/2019/0821.

[69] Àlex Ossó (2019). "New species of Kromtitis Müller, 1984 (Decapoda: Brachyura: Dynomenidae) from the Eocene of Iberian Peninsula". Palaeontologia Electronica. 22 (2): Article number 22.2.47. doi:10.26879/967.

[70] Alessandro Garassino; Patricia G. Weaver; Roger W. Portell; Francisco J. Vega (2019). "A new homolid crab, Lindahomola longispina n. gen., n. sp. (Crustacea, Decapoda), from the Peedee Formation (late Maastrichtian), Rocky Point Member, North Carolina, USA". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 292 (3): 247–258. doi:10.1127/njgpa/2019/0819.

[71] Matúš Hyžný; Herbert Summesberger (2019). "A new species of Mesostylus (Decapoda, Axiidea, Callianassidae) – a peep into the private life of a Late Cretaceous burrowing shrimp". Cretaceous Research. 101: 108–123. doi:10.1016/j.cretres.2019.04.004.

[72] Loris Ceccon; Antonio De Angeli (2019). "Parthenopoidea (Decapoda, Brachyura) dell'Eocene inferiore dei Monti Lessini orientali (Vicenza, Italia nordorientale)". Lavori – Società Veneziana di Scienze Naturali. 44: 85–92.

[PE221Gasparicetal-73] Rok Gašparič; Matúš Hyžný; Gordana Jovanović; Stjepan Ćorić; Sejfudin Vrabac (2019). "Middle Miocene decapod crustacean assemblage from the Tuzla Basin (Tušanj, Bosnia and Herzegovina), with a description of two new species and comparison with coeval faunas from Slovenia" (PDF). Palaeontologia Electronica. 22 (1): Article number 22.1.9. doi:10.26879/894. Archived from the original (PDF) on 2019-04-30.

[74] Francisco J. Vega; Niel L Bruce; Oscar González-León; Jason Jeremiah; María de Lourdes Serrano-Sánchez; Jesús Alvarado-Ortega; Josep A. Moreno-Bedmar (2019). "Lower Cretaceous marine isopods (Isopoda: Cirolanidae, Sphaeromatidae) from the San Juan Raya and Tlayúa formations, Puebla, Mexico". Journal of Crustacean Biology. 39 (2): 121–135. doi:10.1093/jcbiol/ruy107.

[75] Stefan Polkowsky; René H.B. Fraaije (2019). "A new Oligocene hermit crab (Decapoda, Anomura, Paguroidea) from the erratic 'Sternberger Gestein', northern Germany". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 291 (1): 61–63. doi:10.1127/njgpa/2019/0789.

[76] Bao-Jie Du; Rui Chen; Xin-Zheng Li; Wen-Tao Tao; Wen-Jun Bu; Jin-Hua Xiao; Da-Wei Huang (2019). "The first amber caridean shrimp from Mexico reveals the ancient adaptation of the Palaemon to the mangrove estuary environment". Scientific Reports. 9 (1): Article number 14782. Bibcode:2019NatSR...914782D. doi:10.1038/s41598-019-51218-5. PMC 6820862. PMID 31664070.

[77] Loris Ceccon; Antonio de Angeli (2019). "Un nuovo crostaceo Percnidae dell'Oligocene di Sant'Urbano, Vicenza, Italia settentrionale" (PDF). Bollettino del Museo Civico di Storia Naturale di Verona. 43: 17–22.

[Mizunami45Planoprosopon-78] Hiroaki Karasawa; Takayoshi Hirota (2019). "Two new species of Planoprosopon (Decapoda: Brachyura: Longodromitidae) from the Torinosu Group (Upper Jurassic–Lower Cretaceous), Shikoku, Japan" (PDF). Bulletin of the Mizunami Fossil Museum. 45: 27–32. Archived from the original (PDF) on 2019-04-30.

[79] Günter Schweigert (2019). "An additional representative of Lecythocaridae (Decapoda: Brachyura: Dromiacea) from the Upper Jurassic of southern Germany". Journal of Crustacean Biology. 39 (1): 36–39. doi:10.1093/jcbiol/ruy092.

[80] Mario Schädel; Vincent Perrichot; Joachim T. Haug (2019). "Exceptionally preserved cryptoniscium larvae - morphological details of rare isopod crustaceans from French Cretaceous Vendean amber". Palaeontologia Electronica. 22 (3): Article number 22.3.71. doi:10.26879/977.

[81] Krzysztof Hryniewicz; Kazutaka Amano; Maria Aleksandra Bitner; Jonas Hagström; Steffen Kiel; Adiël A. Klompmaker; Thomas Mörs; Cristina M. Robins; Andrzej Kaim (2019). "A late Paleocene fauna from shallow-water chemosynthesis-based ecosystems, Spitsbergen, Svalbard" (PDF). Acta Palaeontologica Polonica. 64 (1): 101–141. doi:10.4202/app.00554.2018. Archived from the original (PDF) on 2019-05-01.

[82] Cristiano Larghi; Andrea Tintori; Daniela Basso; Gianluca Danini; Markus Felber (2019). "A new Ladinian (Middle Triassic) mysidacean shrimp (Crustacea, Lophogastrida) from northern Italy and southern Switzerland". Journal of Paleontology. 94 (2): 291–303. doi:10.1017/jpa.2019.75. S2CID 210618353.

[ChaudharyNagori2019-83] Maya Chaudhary; Madan Lal Nagori (2019). "Ostracods from the Bagh Formation (Upper Cretaceous) of the Narmada Basin, India: Their age and paleobiogeographical significance". Micropaleontology. 65 (6): 513–543.

[PJ532ostracods-84] L.M. Melnikova (2019). "Ostracods from the Delingde Formation (Upper Ordovician) of the Siberian Platform". Paleontological Journal. 53 (2): 140–154. doi:10.1134/S0031030119020072. S2CID 195301704.

[MPS173655-85] I. J. Slipper (2019). "Ostracoda from the Turonian of South-east England". Monographs of the Palaeontographical Society. 173 (655): 1–75. doi:10.1080/02693445.2019.1687983. S2CID 214277821.

[86] Marco Spadi; Elsa Gliozzi; Ian Boomer; Marius Stoica; John Athersuch (2019). "Taxonomic harmonization of Neogene and Quaternary candonid genera (Crustacea, Ostracoda) of the Paratethys". Journal of Systematic Palaeontology. 17 (19): 1665–1698. doi:10.1080/14772019.2018.1545708. S2CID 91396834.

[Monograph172651-87] Vincent Perrier; David J. Siveter; Mark Williams; Douglas Palmer (2019). "British Silurian myodocope ostracods". Monographs of the Palaeontographical Society. 172 (651): 1–99. doi:10.1080/02693445.2018.1509567. S2CID 135217747.

[JSPBergueetal-88] Cristianini Trescastro Bergue; Simone Nunes Brandão; Geise de Santana dos Anjos Zerfass (2019). "Palaeoceanographical events from the late Miocene to Pleistocene of the Rio Grande Rise (south-western Atlantic) as indicated by Ostracoda". Journal of Systematic Palaeontology. 17 (17): 1497–1518. doi:10.1080/14772019.2018.1536895. S2CID 91946254.

[TanakaetalFukujiFm-89] Gengo Tanaka; David J. Siveter; Mark Williams (2019). "Devonian shallow marine ostracods from central Japan". Island Arc. 28 (1): e12283. doi:10.1111/iar.12283.

[BSGF190-90] Marie-Béatrice Forel; Ben Thuy; Max Wisshak (2019). "Digging into the ancestral stocks of Jurassic lineages: ostracods (Crustacea) from Carnian (Late Triassic) sponge mounds from the Maantang Formation (South China)". BSGF - Earth Sciences Bulletin. 190: Article number 9. doi:10.1051/bsgf/2019009.

[Pauproles-91] David J. Siveter; Gengo Tanaka; Mark Williams; Peep Männik (2019). "Japan's earliest ostracods". Island Arc. 28 (1): e12284. doi:10.1111/iar.12284.

[Gibbosocythere-92] Maria Karpuk; Robin Whatley; Caroline Maybury (2019). "New marine Ostracoda from the Early Cretaceous (upper Barremian – Aptian) of the Crimean Peninsula". Revue de Micropaléontologie. 62 (1): 1–8. doi:10.1016/j.revmic.2018.11.002.

[Micropaleontology652Krithe-93] Cristianini Trescastro Bergue; Joao Carlos Coimbra; Kenneth L. Finger (2019). "A new taxonomic approach to Krithe Brady, Crosskey and Robertson, 1874 (Crustacea: Ostracoda) based on specimens from the Miocene of Chile". Micropaleontology. 65 (2): 161–171.

[94] Vincent Perrier; Ewa Olempska; David J. Siveter; Mark Williams; Nicolas Legiot (2019). "Silurian myodocope ostracods from Poland" (PDF). Acta Palaeontologica Polonica. 64 (2): 379–397. doi:10.4202/app.00552.2018. Archived from the original (PDF) on 2019-05-01.

[ChaudharyetalJPSI2019-95] Maya Chaudhary; M.L. Nagori; S. C. Khosla (2019). "Ostracod genus Rostrocytheridea from the Bagh Formation (Upper Cretaceous) of Narmada Basin, Madhya Pradesh and Gujarat, India". Journal of the Palaeontological Society of India. 64 (1): 73–82.

[96] Alberto Collareta; Giovanni Coletti; Giulia Bosio; John Buckeridge; Christian de Muizon; Thomas J. DeVries; Rafael M. Varas-Malca; Alí Altamirano-Sierra; Mario Urbina-Schmitt; Giovanni Bianucci (2019). "A new barnacle (Cirripedia: Neobalanoformes) from the early Miocene of Peru: Palaeoecological and palaeobiogeographical implications". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 292 (3): 321–338. doi:10.1127/njgpa/2019/0825.

[ZLcalmanostracan-97] Philipp Wagner; Joachim T. Haug; Carolin Haug (2019). "A new calmanostracan crustacean species from the Cretaceous Yixian Formation and a simple approach for differentiating fossil tadpole shrimps and their relatives". Zoological Letters. 5: Article 20. doi:10.1186/s40851-019-0136-0. PMC 6582493. PMID 31245037.

[Jaegerscalpellum-98] Andy Gale (2019). "Thoracican cirripedes (Crustacea) from the Hauterivian (Lower Cretaceous) of Hannover, northern Germany". Proceedings of the Geologists' Association. 130 (6): 659–672. doi:10.1016/j.pgeola.2019.03.002.

[99] Andrew Scott Gale (2020). "New thoracican cirripedes (Crustacea) from the Cretaceous of Europe and North Africa". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 295 (3): 243–282. doi:10.1127/njgpa/2020/0886.

[100] Carrie E. Schweitzer; Rodney M. Feldmann; Megan M. Schinker (2019). "A new species of Cyclida (Multicrustacea; Halicynidae) from the Triassic of The Netherlands". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 293 (1): 67–71. doi:10.1127/njgpa/2019/0831.

[101] Oscar F. Gallego; Shen Yan-Bin; Ed Jarzembowski; Ian J. Slipper; Angela Self; Mateo D. Monferran (2019). "The Crustacea of the Insect Bed (latest Eocene) of the Isle of Wight, England, including the first spinicaudatan (clam shrimp) from the British Cenozoic". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 110 (3–4): 289–299. doi:10.1017/S1755691018000452.

[102] Huanyu Liao; Yanbin Shen; Jian Gao; Diying Huang (2019). "Pemphilimnadiopsis cheni sp. nov. (Branchiopoda: Diplostraca: Spinicaudata) from the Upper Carboniferous of East Hebei, China and its biostratigraphic significance". Palaeoentomology. 2 (4): 381–389. doi:10.11646/palaeoentomology.2.4.12.

[103] Giovanni Coletti; Alberto Collareta; Giulia Bosio; Mario Urbina-Schmitt; John Buckeridge (2019). "Perumegabalanus calziai gen. et sp. nov., a new intertidal megabalanine barnacle from the early Miocene of Peru". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 294 (2): 197–212. doi:10.1127/njgpa/2019/0856. hdl:10281/248752.

[104] Alberto Collareta; Agatino Reitano; Antonietta Rosso; Rossana Sanfelippo; Mark Bosselaers; Giovanni Bianucci; Gianni Insacco (2019). "The oldest platylepadid turtle barnacle (Cirripedia, Coronuloidea): a new species of Platylepas from the Lower Pleistocene of Italy". European Journal of Taxonomy. 516 (516): 1–17. doi:10.5852/ejt.2019.516.

[105] Eduard V. Mychko; Rodney M. Feldmann; Carrie E. Schweitzer; Alexander S. Alekseev (2019). "New genus of Cyclida (Crustacea) from Lower Carboniferous (Mississippian, Viséan) of Russia and England and new species from Viséan of Russia". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 294 (1): 81–90. doi:10.1127/njgpa/2019/0847.

[ZT47121-106] Ray T. Perreault; John S. Buckeridge (2019). "Paleogene Verrucidae (Cirripedia: Verrucomorpha) of North America, with descriptions of three new species". Zootaxa. 4712 (1): 34–50. doi:10.11646/zootaxa.4712.1.2. PMID 32230694.

[107] John R. Paterson; Gregory D. Edgecombe; Michael S. Y. Lee (2019). "Trilobite evolutionary rates constrain the duration of the Cambrian explosion". Proceedings of the National Academy of Sciences of the United States of America. 116 (10): 4394–4399. Bibcode:2019PNAS..116.4394P. doi:10.1073/pnas.1819366116. PMC 6410820. PMID 30782836.

[108] Stephen Pates; Russell D.C. Bicknell (2019). "Elongated thoracic spines as potential predatory deterrents in olenelline trilobites from the lower Cambrian of Nevada". Palaeogeography, Palaeoclimatology, Palaeoecology. 516: 295–306. Bibcode:2019PPP...516..295P. doi:10.1016/j.palaeo.2018.12.013.

[109] Guang-Ying Du; Jin Peng; De-Zhi Wang; Qiu-Jun Wang; Yi-Fan Wang; Hui Zhang (2019). "Morphology and developmental traits of the trilobite Changaspis elongata from the Cambrian Series 2 of Guizhou, South China". Acta Palaeontologica Polonica. 64 (4): 797–813. doi:10.4202/app.00604.2019.

[110] David R. Schwimmer; William M. Montante (2019). "A Cambrian meraspid cluster: evidence of trilobite egg deposition in a nest site". PALAIOS. 34 (5): 254–260. Bibcode:2019Palai..34..254S. doi:10.2110/palo.2018.102. S2CID 181784455.

[111] Fernanda Serra; Diego Balseiro; Beatriz G. Waisfeld (2019). "Diversity patterns in upper Cambrian to Lower Ordovician trilobite communities of north‐western Argentina". Palaeontology. 62 (4): 677–695. doi:10.1111/pala.12424.

[112] Daniela S. Monti; Viviana A. Confalonieri (2019). "First cladistic analysis of the trilobite family Olenidae from the Furongian and Ordovician". Lethaia. 52 (3): 304–322. doi:10.1111/let.12313.

[113] Melanie J. Hopkins (2019). "Phylogenetic analysis and revision of the trilobite subfamily Balnibarbiinae (Olenidae)". American Museum Novitates. 2019 (3928): 1–20. doi:10.1206/3928.1. hdl:2246/6938. S2CID 164469838.

[114] Harriet B. Drage; Thijs R.A. Vandenbroucke; Peter Van Roy; Allison C. Daley (2019). "Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco". Acta Palaeontologica Polonica. 64 (2): 261–273. doi:10.4202/app.00582.2018.

[115] Jean Vannier; Muriel Vidal; Robin Marchant; Khadija El Hariri; Khaoula Kouraiss; Bernard Pittet; Abderrazak El Albani; Arnaud Mazurier; Emmanuel Martin (2019). "Collective behaviour in 480-million-year-old trilobite arthropods from Morocco". Scientific Reports. 9 (1): Article number 14941. Bibcode:2019NatSR...914941V. doi:10.1038/s41598-019-51012-3. PMC 6797724. PMID 31624280.

[116] Morgane Oudot; Catherine Crônier; Pascal Neige; David Holloway (2019). "Phylogeny of some Devonian trilobites and consequences for the systematics of Austerops (Phacopidae)". Journal of Systematic Palaeontology. 17 (9): 775–790. doi:10.1080/14772019.2018.1471105. S2CID 91163530.

[117] Gabriel S. Jacobs; Jesse R. Carlucci (2019). "Ontogeny and shape change of the phacopid trilobite Calyptaulax". Journal of Paleontology. 93 (6): 1105–1125. doi:10.1017/jpa.2019.38. S2CID 199098897.

[118] Russell D.C. Bicknell; John R. Paterson; Melanie J. Hopkins (2019). "A trilobite cluster from the Silurian Rochester Shale of New York: predation patterns and possible defensive behavior". American Museum Novitates. 2019 (3937): 1–16. doi:10.1206/3937.1. hdl:2246/6959. S2CID 202401576.

[119] Morgane Oudot; Pascal Neige; Rémi Laffont; Nicolas Navarro; Ahmed Yacine Khaldi; Catherine Crônier (2019). "Functional integration for enrolment constrains evolutionary variation of phacopid trilobites despite developmental modularity". Palaeontology. 62 (5): 805–821. doi:10.1111/pala.12428.

[120] Gerhard Scholtz; Andreas Staude; Jason A. Dunlop (2019). "Trilobite compound eyes with crystalline cones and rhabdoms show mandibulate affinities". Nature Communications. 10 (1): Article number 2503. Bibcode:2019NatCo..10.2503S. doi:10.1038/s41467-019-10459-8. PMC 6555793. PMID 31175282.

[121] Harriet B. Drage (2019). "Quantifying intra- and interspecific variability in trilobite moulting behaviour across the Palaeozoic". Palaeontologia Electronica. 22 (2): Article number 22.2.34. doi:10.26879/940.

[122] Xin Wei; Renbin Zhan (2019). "A new trilobite species from the upper Rhuddanian (lower Silurian) of South China, and its biogeographical implications". Alcheringa: An Australasian Journal of Palaeontology. 43 (1): 33–42. doi:10.1080/03115518.2018.1508614. S2CID 134818599.

[trilobitesDehMolla-123] Mansoureh Ghobadi Pour (2019). "Ordovician trilobites from Deh-Molla, eastern Alborz, Iran". Alcheringa: An Australasian Journal of Palaeontology. 43 (3): 381–405. doi:10.1080/03115518.2019.1616110. S2CID 218636212.

[GB2212trilobites-124] Allart P. van Viersen; Peter Taghon; Benedikt Magrean (2019). "Early Middle Devonian trilobites and events in the Nismes – Vireux-Molhain area, southern border of the Dinant Synclinorium (Belgium, northern France)". Geologica Belgica. 22 (1–2): 7–33. doi:10.20341/gb.2019.001.

[125] Michal Mergl; Petr Budil (2019). "Rhynchonelliform brachiopods and trilobites of the 'upper dark interval' in the Koněprusy area (Devonian, Eifelian, Kačák Event; the Czech Republic)". Fossil Imprint. 75 (1): 92–107. doi:10.2478/if-2019-0008. S2CID 203606755.

[FossilsStrata64-126] Gerd Geyer (2019). "The earliest known West Gondwanan trilobites from the Anti‐Atlas of Morocco, with a revision of the family Bigotinidae Hupe, 1953". Fossils and Strata. 64: 55–154. doi:10.1002/9781119564249.ch6. ISBN 978-1-119-56423-2.

[127] Eduard V. Mychko; Oles V. Savchuk (2019). "A new brachymetopid trilobite from the Early Permian Shakhtau reef complex of the southwestern Urals, Bashkortostan, Russia". Zootaxa. 4555 (3): 346–358. doi:10.11646/zootaxa.4555.3.4. PMID 30790921.

[MuellerHahn-128] Peter Müller; Gerhard Hahn (2019). "A new trilobite frauna from the Mississippian of Rösenbeck (North Rhine-Westphalia, Germany)". Dortmunder Beiträge zur Landeskunde, Naturwissenschaftliche Mitteilungen. 49: 93–161.

[JSPOveEbbestadFortey-129] Jan Ove; R. Ebbestad; Richard A. Fortey (2020). "Late Ordovician trilobites from the Taimyr Peninsula, Arctic Russia". Journal of Systematic Palaeontology. 18 (1): 1–135. doi:10.1080/14772019.2019.1601643. S2CID 201206166.

[JPCyphaspides-130] Brian D.E. Chatterton; Stacey Gibb; Ryan C. McKellar (2019). "Species of the Devonian aulacopleurid trilobite Cyphaspides from southeastern Morocco". Journal of Paleontology. 94 (1): 99–114. doi:10.1017/jpa.2019.71. S2CID 204269182.

[MGM47Alberti-131] Mirco Alberti (2019). "Die Trilobiten Cyphaspis konstantini n. sp. (Aulacopleuridae) und Koneprusia chimaera n. sp. (Odontopleuridae) aus dem basalen Rupbach-Schiefer (Ober-Emsium; Rhenoherzyniklum)". Mainzer Geowissenschaftliche Mitteilungen. 47: 27–54.

[Cyrtosymbolinella-132] Martin Basse; Ulrich Lemke (2019). "New and little known trilobites from the Early and Late Devonian of the Kellerwald (Rhenohercynian Zone, German Variscides)". Dortmunder Beiträge zur Landeskunde, Naturwissenschaftliche Mitteilungen. 49: 47–69.

[133] Martin Basse; Peter Müller; Jörg Habenicht (2019). "Heliopeltis and Sagittapeltis in the Emsian of the Rhenohercynian Zone (Trilobita, Early Devonian, German Variscides)". Dortmunder Beiträge zur Landeskunde, Naturwissenschaftliche Mitteilungen. 49: 71–91.

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