2012 in fish paleontology

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This list of fossil fish species is a list of taxa of fish that have been described during the year 2012. The list only includes taxa at the level of genus or species.

Newly named jawless vertebrates

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Diademaspis janvieri[1]	Sp. nov	Valid	Keating, Sansom & Purnell	Early Devonian (Lochkovian)	Wayne Herbert Quarry	United Kingdom	A member of Osteostraci, a species of Diademapsis.
Dunyu[2]	Gen. et sp. et comb. nov	Valid	Zhu, Liu, Jia & Gai	Silurian	Huixingshao Formation Kuanti Formation	China	A eugaleaspidiform galeaspid. Its type species is Dunyu longiforus; genus also contains "Eugaleaspis" xiushanensis Liu 1983.
Janaspis[1]	Gen. et comb et 2 sp. nov	Valid	Keating, Sansom & Purnell	Early Devonian (Lochkovian)	Wayne Herbert Quarry	United Kingdom	A new genus of Osteostraci including the type species "Cephalaspis" pagei Lankester, 1868[3] and new species J. newtonensis and J. punctata.
Kerreralepis[4]	Gen. et sp. nov	Valid	Blom	Lower Devonian		United Kingdom	A birkeniid anaspid. The type species is Kerreralepis carinata.
Waengsjoeaspis platycornis[5]	Sp. nov	Valid	Scott & Wilson	Early Devonian (Lochkovian)	Man On The Hill Lagerstätte	Canada	A member of Osteostraci, a species of Waengsjoeaspis.
Zenaspis waynensis[1]	Sp. nov	Valid	Keating, Sansom & Purnell	Early Devonian (Lochkovian)	Wayne Herbert Quarry	United Kingdom	A member of Osteostraci, a species of Zenaspis.

Newly named acanthodians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Halimacanthodes[6]	Gen. et sp. nov	Valid	Burrow, Trinajstic & Long	Late Devonian (Frasnian)	Gogo Formation	Australia	An acanthodiform acanthodian. The type species is Halimacanthodes ahlbergi.
Machaeracanthus goujeti[7]	Sp. nov	Valid	Botella, Martínez-Pérez & Soler-Gijón	Early Devonian		France Spain	A species of Machaeracanthus.
Podoliacanthus[8]	Gen. et sp. nov	Valid	Voichyshyn & Szaniawski	Early Devonian, possibly also late Silurian.		Ukraine Greenland	An ischnacanthiform, possibly an ischnacanthid. The type species is Podoliacanthus zychi; also contains three yet unnamed species.

Newly named cartilaginous fishes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Abdounia lata[9]	Sp. nov	Valid	Malyshkina	Late Eocene	Tavda Formation	Russia	A requiem shark, a species of Abdounia.
Abdounia vassilyevae[9]	Sp. nov	Valid	Malyshkina	Late Eocene	Tavda Formation	Russia	A requiem shark, a species of Abdounia.
Agaleorhynchus[10]	Gen. et sp. nov	Valid	Guinot et al.	Late Cretaceous (late Santonian to early Campanian)		United Kingdom	A sclerorhynchid. The type species is Agaleorhynchus britannicus.
Argoubia[11]	Gen. et sp. nov	Valid	Adnet et al.	Late Eocene		Morocco	A mobulid. The type species is Argoubia barbei.
Australopristis[12]	Gen. et sp. nov	Valid	Martill & Ibrahim	Late Cretaceous		New Zealand	A sawfish. The type species is Australopristis wiffeni.
Borodinopristis shannoni[13]	Sp. nov	Valid	Case et al.	Late Cretaceous (Campanian)		United States	A sclerorhynchid sawfish, a species of Borodinopristis.
Carcharodon hubbelli[14]	Sp. nov	Valid	Ehret et al.	Late Miocene	Pisco Formation	Peru	A lamnid shark, a species of Carcharodon.
Cretaplatyrhinoidis[10]	Gen. et sp. nov	Valid	Guinot et al.	Late Cretaceous (Santonian to Campanian)		France United Kingdom	A platyrhinid. The type species is Cretaplatyrhinoidis ornatus.
Cromhallia[15]	Gen. et sp. nov	Valid	Behan, Walken & Cuny	Early Carboniferous		United Kingdom	A chondrichthyan of indeterminate phylogenetic relationships. The type species is Cromhallia parvunda.
Duffinselache[16]	Gen. et comb. nov	Valid	Andreev & Cuny	Rhaetian		United Kingdom	A selachimorph, a new genus for "Polyacrodus" holwellensis (Duffin, 1998).
Eoplinthicus underwoodi[11]	Sp. nov	Valid	Adnet et al.	Late Eocene		Morocco	A mobulid, a species of Eoplinthicus.
Fragilicorona wanlantricuspis[17]	Sp. nov	Valid	Zhang et al.	Middle Triassic (Ladinian)	Zhuganpo Formation	China	An elasmobranch, a species of Fragilicorona.
Galeocorax[18]	Nom. nov	Valid	Cappetta	Late Cretaceous		United Kingdom United States	A shark related to Pseudocorax;[19] a replacement name for Paracorax Cappetta (1977).
Glabrisubcorona triridgecorona[17]	Sp. nov	Valid	Zhang et al.	Middle Triassic (Ladinian)	Zhuganpo Formation	China	An elasmobranch, a species of Glabrisubcorona.
Isacrodus[20]	Gen. et sp. nov	Valid	Ivanov, Nestell & Nestell	Middle Permian (Roadian)	Cutoff Formation	United States	A jalodontid chondrichthyan. The type species is Isacrodus marthae.
Kaibabvenator[21]	Gen. et sp. nov	Valid	Hodnett et al.	Permian (Late Leonardian)	Kaibab Limestone	United States	A ctenacanthiform. The type species is Kaibabvenator swiftae.
Nanocorax[18]	Nom. nov	Valid	Cappetta	Late Cretaceous		France United Kingdom United States	A member of Anacoracidae; a replacement name for Microcorax Cappetta & Case (1975).
Nanoskalme[21]	Gen. et sp. nov	Valid	Hodnett et al.	Permian (Late Leonardian)	Kaibab Limestone	United States	A ctenacanthiform. The type species is Nanoskalme natans.
Neosaivodus[21]	Gen. et sp. nov	Valid	Hodnett et al.	Permian (Late Leonardian)	Kaibab Limestone	United States	A ctenacanthiform. The type species is Neosaivodus flagstaffensis.
Odontaspis watinensis[22]	Sp nov	Valid[23]	Cook et al.	Late Cretaceous (early Turonian)	Kaskapau Formation	Canada	A species of Odontaspis.
Oromobula[11]	Gen. et sp. nov	Valid	Adnet et al.	Late Eocene		Morocco	A mobulid. The type species is Oromobula dakhlaensis.
Parasquatina jarvisi[24]	Sp nov	Valid	Guinot et al.	Late Cretaceous (late Santonian to early late Campanian)		France United Kingdom	A squatiniform, a species of Parasquatina.
Parasquatina justinensis[24]	Sp nov	Valid	Guinot et al.	Late Cretaceous (middle Turonian)		France	A squatiniform, a species of Parasquatina.
Plesiobatis vandenboschi[25]	Sp. nov	Valid	Bor, Reinecke & Verschueren	Miocene (early Langhian)	Breda Formation	Netherlands	A relative of the deepwater stingray.
Proteothrinax[26]	Nom. nov	Valid	Pfeil	Eocene (Lutetian)	Weitwies Subformation	Austria	A member of Chlamydoselachidae; a replacement name for Thrinax Pfeil (1983).
Pseudoplatyrhina[10]	Gen. et sp. nov	Valid	Guinot et al.	Late Cretaceous (late Santonian to early Campanian)		France United Kingdom	A platyrhinid. The type species is Pseudoplatyrhina crispa.
Ptychotrygonoides sabatieri[10]	Sp. nov	Valid	Guinot et al.	Late Cretaceous (middle Turonian)		France	A ptychotrygonid, possibly a relative of sclerorhynchids; a species of Ptychotrygonoides.
"Rhinobatos" seruensis[10]	Sp. nov	Valid	Guinot et al.	Late Cretaceous (late Santonian)		France	A guitarfish, a possible species of Rhinobatos.
Rhomaleodus[16]	Gen. et sp. nov	Valid	Andreev & Cuny	Anisian		Bulgaria	A selachimorph. The type species is Rhomaleodus budurovi.
Rugosicorona circacarina[17]	Sp. nov	Valid	Zhang et al.	Middle Triassic (Ladinian)	Zhuganpo Formation	China	An elasmobranch, a species of Rugosicorona.
Sphenacanthus riorastoensis[27]	Sp. nov	Valid	Pauliv, Dias & Sedor	Permian (Wordian to Wuchiapingian)	Rio do Rasto Formation	Brazil	A species of Sphenacanthus.
Squatirhina draytoni[10]	Sp. nov	Valid	Guinot et al.	Late Cretaceous (late Early Cenomanian to middle Turonian)		France United Kingdom	A guitarfish, a species of Squatirhina.
Tantalepis[28]	Gen. et sp. nov	Valid	Sansom et al.	Middle Ordovician (Darriwilian)	Stairway Sandstone	Australia	A gnathostome of uncertain phylogenetic placement, a possible cartilaginous fish. The type species is Tantalepis gatehousei.
Texasodus[20]	Gen. et sp. nov	Valid	Ivanov, Nestell & Nestell	Middle Permian (late Wordian or early Capitanian)	Bell Canyon Formation	USA	A jalodontid chondrichthyan. The type species is Texasodus varidentatus.

Newly named bony fishes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Aglyptorhynchus hakataramea[29]	sp nov	Valid	Gottfried, Fordyce & Rust	Late Oligocene		New Zealand	A billfish
Ampheristus neobavaricus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A cusk-eel, a species of Ampheristus.
Anguilla pfeili[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian)	Oiching Formation	Germany	A member of Anguillidae, a species of Anguilla.
Apsopelix miyazakii[31]	Sp. nov	Valid	Yabumoto, Hikida & Nishino	Late Cretaceous (Turonian)		Japan	A species of Apsopelix.
Argyripnus kroisbachensis[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A member of Sternoptychidae, a species of Argyripnus.
Aspidopleurus kickapoo[32]	Sp. nov	Valid	Alvarado-Ortega & Porras-Múzquiz	Late Cretaceous (Turonian)	Eagle Ford Formation	Mexico	An aspidopleurid aulopiform, a species of Aspidopleurus.
Aspistor verumquadriscutis[33]	sp nov	Valid	Aguilera & Marceniuk	Late Miocene	Urumaco Formation	Venezuela	An ariid, a species of Aspistor.
Aulopus praeteritus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A member of Aulopidae, a species of Aulopus.
Bavariscopelus parvinavis[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian)	Oiching Formation	Germany	A member of Myctophiformes of uncertain phylogenetic placement, a species of Bavariscopelus.
Bawitius[34]	Gen. et comb. nov	Valid	Grandstaff et al.	Cenomanian	Bahariya Formation	Egypt	A polypterid actinopterygian, a new genus for "Polypterus" bartheli (Schaal, 1984).
Belonostomus marquesbritoi[35]	Sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Campanian-Maastrichtian boundary)		Italy	A member of Aspidorhynchiformes, a species of Belonostomus.
Bruehnopteron[36]	Gen. et sp. nov	Valid	Schultze & Reed	Middle Devonian		United States	A tristichopterid sarcopterygian. The type species is Bruehnopteron murphyi. The species was argued by Parfitt et al. (2014) to be likely junior synonym of Tinirau clackae.[37]
Caruso[38]	Gen. et comb. nov	Valid	Carnevale & Pietsch	Eocene		Italy	A lophiid anglerfish, a new genus for "Lophius" brachysomus (Agassiz).
Centroberyx apogoniformis[30]	Sp. nov	Valid	Schwarzhans	Late Cretaceous (Maastrichtian)[39] and Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany United States ( Arkansas Texas)[39]	A member of Berycidae, a species of Centroberyx.
Cyclogonostoma[30]	Gen. et sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A member of Gonostomatidae. The type species is Cyclogonostoma disciformis.
Danoscopelus[30]	Gen. et comb. nov	Valid	Schwarzhans	Paleocene (Selandian to Thanetian)	Oiching Formation	Austria Denmark	A member of Myctophiformes of uncertain phylogenetic placement; a new genus for "genus Myctophidarum" schnetleri Schwarzhans (2003).
Dentex? solidus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A member of Sparidae, possibly a species of Dentex.
Deutschenchelys[40]	Gen. et sp. nov	Valid	Prokofiev	Early Oligocene		Germany	An eel, possibly a member of Moringuidae. The type species is Deutschenchelys micklichi.
Diandongperleidus[41]	Gen. et sp. nov	Valid	Geng et al.	Middle Triassic (middle or late Anisian)	Guanling Formation	China	A perleidid perleidiform. The type species is Diandongperleidus denticulatus.
Diretmus serrativenter[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A spinyfin related to the silver spinyfin.
Echinelops[42]	Gen. et sp. nov.	Valid	Murray & Hoşgör	Early Oligocene		Turkey	An elopiform fish. The type species is Echinelops ozcani.
Eotetraodon tavernei[43]	Sp. nov	Valid	Tyler & Bannikov	Eocene (Ypresian or Lutetian)		Italy	A pufferfish, a species of Eotetraodon.
Eurolebias[44]	Gen. et comb. nov	Valid	Costa	Late Oligocene		France	A pupfish; a new genus for "Prolebias" meridionalis Gaudant (1978).
Francolebias[44]	Gen. et comb. nov	Valid	Costa	Early Oligocene		France	A relative of Valencia; a new genus for "Prolebias" delphinensis Gaudant (1989) and "Lebias" aymardi Sauvage (1869).
Frigosorbinia[45]	Gen. et comb. nov	Valid	Bannikov & Tyler	Eocene		Italy	A surgeonfish; a new genus for "Pesciarichthys" baldwinae Sorbini & Tyler (1998).
Frippia[46]	Gen. et sp. nov	Valid	Bannikov & Carnevale	Eocene		Italy	A member of Percoidei. The type species is Frippia labroiformis.
Fuyuanperleidus[41]	Gen. et sp. nov	Valid	Geng et al.	Middle Triassic (late Ladinian)	Falang Formation	China	A perleidid perleidiform. The type species is Fuyuanperleidus dengi.
Galaxias angustiventris[47]	Sp nov	Valid	Schwarzhans in Schwarzhans et al.	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias brevicauda[47]	Sp nov	Valid	Schwarzhans in Schwarzhans et al.	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias bobmcdowalli[47]	Sp nov	Valid	Schwarzhans in Schwarzhans et al.	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias papilionis[47]	Sp nov	Valid	Schwarzhans in Schwarzhans et al.	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias parvirostris[47]	Sp nov	Valid	Schwarzhans in Schwarzhans et al.	Early Miocene	Bannockburn Formation	New Zealand	A galaxiid, a species of Galaxias.
Galaxias tabidus[47]	Sp nov	Valid	Schwarzhans in Schwarzhans et al.	Early Miocene		New Zealand	A galaxiid, a species of Galaxias.
Gerpegezhus[48]	Gen. et sp. nov	Valid	Bannikov & Carnevale	Early Eocene		Russia	A close relative of centriscids. The type species is Gerpegezhus paviai.
Gnathophis probus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A member of Congridae, a species of Gnathophis.
Gorgasia? turgidus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A member of Congridae, possibly a species of Gorgasia.
Haemulon? conjugator[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Selandian)	Oiching Formation	Austria Germany	A member of Haemulidae of uncertain phylogenetic placement.
Hippotropiscis[49]	Gen. et sp. nov	Valid	Žalohar & Hitij	Middle Miocene		Slovenia	A hippocampine syngnathid. The type species is Hippotropiscis frenki.
Holocentronotus blandus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A member of Holocentridae, a species of Holocentronotus.
Ictalurus countermani[50]	sp nov	Valid	Lundberg & Luckenbill	Late Miocene	St. Marys Formation	United States	A species of Ictalurus.
Illusionella[51]	Gen. et 2 sp. nov	Valid	Baykina	Miocene		Russia	A clupeid fish. Genus contains two species: Illusionella tsurevica and Illusionella pshekhensis. The genus Illusionella was subsequently synonymized with the genus Sarmatella Menner (1949) by Baykina (2013); however, Baykina kept the species I. tsurevica and I. pshekhensis as valid, distinct species within the genus Sarmatella.[52]
Isozen mareikeae[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A member of Zeiformes of uncertain phylogenetic placement, related to the family Parazenidae; a species of Isozen.
Isurichthys breviusculus[53]	Sp nov	Valid	Bannikov	Lower Oligocene		Russia	An ariommatid, a species of Isurichthys.
Kressenbergichthys[30]	Gen. et sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A member of Berycidae. The type species is Kressenbergichthys kuhni.
Kyphosichthys[54]	Gen. et sp. nov	Valid	Xu & Wu	Middle Anisian	Guanling Formation	China	A ginglymodian actinopterygian.
Lactarius? simplex[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A relative of the false trevally.
Lebonichthys nardoensis[55]	Sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Campanian-Maastrichtian boundary)		Italy	A bonefish, a species of Lebonichthys.
Luopingperleidus[41]	Gen. et sp. nov	Valid	Geng et al.	Middle Triassic (middle or late Anisian)	Guanling Formation	China	A perleidid perleidiform. The type species is Luopingperleidus sui.
Luoxiongichthys[56]	Gen. et sp. nov	Valid	Wen et al.	Middle Anisian	Guanling Formation	China	A basal actinopterygian.
Macrosemimimus[57]	Gen. et sp. et comb. nov	Valid	Schröder, López-Arbarello & Ebert	Late Jurassic		France Germany United Kingdom	A semionotiform. The type species is Macrosemimimus fegerti; "Lepidotes" lennieri (Sauvage, 1893) is a second species of Macrosemimimus.
Mataichthys[47]	Gen. et sp. nov	Valid	Schwarzhans, Scofield, Tennyson & T. Worthy in Schwarzhans et al.	Early Miocene	Bannockburn Formation	New Zealand	An eleotrid. Contains M. bictenatus, M. procerus, M. rhinoceros, and M. taurinus
Melamphaes? protoforma[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A ridgehead of uncertain phylogenetic placement.
Mercediella[58]	Nom. nov	Valid	Koerber	Early Cretaceous	Riachuelo Formation	Brazil	A member of Pycnodontiformes; a replacement name for Camposichthys Figueiredo & Silva Santos (1991).
Nanaichthys[59]	Gen. et sp. nov	Valid	Amaral & Brito	Early Cretaceous, possibly Aptian	Marizal Formation	Brazil	A member of Chanidae. The type species is Nanaichthys longipinnus.
Nardoelops[55]	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Campanian-Maastrichtian boundary)		Italy	A member of Elopidae. The type species is Nardoelops nybelini.
Nettastoma davejohnsoni[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Selandian to Thanetian)	Oiching Formation	Austria	A member of Nettastomatidae, a species of Nettastoma.
Ogcocephalus? semen[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A member of Ogcocephalidae, possibly a species of Ogcocephalus.
Palaeogadus? bratishkoi[30]	Sp. nov	Valid	Schwarzhans	Late Cretaceous (Maastrichtian)[39] and Paleocene (Danian[39] and Thanetian)	Kemp Clay Oiching Formation	Austria Denmark[39] United States ( Texas)[39]	Originally described as a member of Merlucciidae and a possible species of Palaeogadus. Subsequently reinterpreted as a stem gadiform of uncertain familial affinities and transferred to the genus Archaemacruroides by Schwarzhans & Stringer (2020).[39]
Palatinichthys[60]	Gen. et sp. nov	Valid	Witzmann & Schoch	Lower Permian	Saar–Nahe Basin	Germany	A megalichthyid sarcopterygian. The type species is Palatinichthys laticeps.
Paleotetra[61]	Gen. et 2 sp.	Valid	Weiss, Malabarba & Malabarba	Eocene to Oligocene		Brazil	A characiform, probably a characid. The genus contains two species: Paleotetra entrecorregos and Paleotetra aiuruoca.
Paraconger vetustus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian)	Oiching Formation	Germany	A member of Congridae, a species of Paraconger.
Paraulopus novellus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian, possibly Thanetian)	Oiching Formation	Germany Austria?	A species of Paraulopus.
Paranaichthys[62]	Gen. et sp. nov	Valid	Dias	Permian, Guadalupian (Wordian or Capitanian)	Rio do Rasto Formation	Brazil	A probable relative of Platysomus and other platysomids. The type species is Paranaichthys longianalis.
Parawenzichthys[63]	Gen. et sp. nov	Valid	de Figueiredo, Gallo & Delarmelina	Turonian	Atlântida Formation	Brazil	A protacanthopterygian.
Platysepta kressenbergensis[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A member of Sparidae, a species of Platysepta.
Plesiopoma elegantissima[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian)	Oiching Formation	Germany	A member of Acropomatidae, a species of Plesiopoma.
Plesiopoma traubi[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A member of Acropomatidae, a species of Plesiopoma.
Polymixia polita[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A beardfish.
Polyperca exserta[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian)	Oiching Formation	Germany	A member of Serranidae, a species of Polyperca.
Progonostoma[30]	Gen. et 2 sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A member of Gonostomatidae. The type species is Progonostoma primordialis; genus also contains Progonostoma hagni.
Prototroctes modestus[47]	Sp nov	Valid	Schwarzhans in Schwarzhans et al.	Early Miocene	Bannockburn Formation	New Zealand	A retropinnid, a species of Prototroctes.
Prototroctes vertex[47]	Sp nov	Valid	Schwarzhans in Schwarzhans et al.	Early Miocene	Bannockburn Formation	New Zealand	A retropinnid, a species of Prototroctes.
Rebellatrix[64]	Gen. et sp. nov	Valid	Wendruff & Wilson	Lower Triassic	Sulphur Mountain Formation	Canada	A coelacanth. The type species is Rebellatrix divaricerca.
Reidus[65]	Gen. et sp. nov	Valid	Graf	Albian		United States	A coelacanth. The type species is Reidus hilli.
Rhinodipterus kimberleyensis[66]	Sp nov	Valid	Clement	Late Devonian	Gogo Formation	Australia	A lungfish, a species of Rhinodipterus.
Rhynchoconger intercedens[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A member of Congridae, a species of Rhynchoconger.
Sangiorgioichthys valmarensis[67]	Sp. nov	Valid	Lombardo, Tintori & Tona	Middle Triassic (late Ladinian)		Switzerland	A semionotid semionotiform, a species of Sangiorgioichthys.
Schaefferius[68]	Nom. nov	Valid	Akbulut & Özdikmen	Early Triassic		Angola	A member of Palaeonisciformes belonging to the family Canobiidae; a replacement name for Marquesia Schaeffer (1990).
Sinamia liaoningensis[69]	sp nov	Valid	Zhang	Early Cretaceous (Barremian to Aptian)	Jiufotang Formation Yixian Formation	China	A sinamiid amiiform, a species of Sinamia.
Squamibolcoides[70]	Gen. et sp. nov	Valid	Bannikov & Zorzin	Eocene		Italy	A member of Percoidei of uncertain phylogenetic placement. The type species is Squamibolcoides minciottii.
Thoracopterus wushaensis[71]	Sp. nov	Valid	Tintori et al.	Middle Triassic (late Ladinian)	Falang Formation	China	A thoracopterid peltopleuriform, a species of Thoracopterus. The species might be synonymous with the species Potanichthys xingyiensis Xu et al. (2013).[72]
Tinirau[73]	Gen. et sp. nov	Valid	Swartz	Middle Devonian	Red Hill I beds	United States	An eotetrapodiform tetrapodomorph related to the clade Elpistostegalia[73] or a tristichopterid.[37] The type species is Tinirau clackae.
Tomognathus gigeri[74]	sp nov	Valid	Cavin & Giner	Valanginian		France	A halecomorph fish, a species of Tomognathus.
Trachichthys anomalopsoides[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A slimehead, a species of Trachichthys.
Trachichthys impavidus[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Danian to Thanetian)	Oiching Formation	Austria Germany	A slimehead, a species of Trachichthys.
Tungsenia[75]	Gen. et sp. nov	Valid	Lu et al.	Early Devonian		China	A basal member of Tetrapodomorpha. The type species is Tungsenia paradoxa.
Valenciennellus kennetti[30]	Sp. nov	Valid	Schwarzhans	Paleocene (Thanetian)	Oiching Formation	Austria	A member of Sternoptychidae, a species of Valenciennellus.
Voelklichthys[76]	Gen. et sp. nov	Valid	Arratia & Schultze	Kimmeridgian		Germany	A macrosemiiform. The type species is Voelklichthys comitatus.
Zoqueichthys[77]	Gen. et sp. nov	Valid	Alvarado-Ortega & Than-Marchese	Late Cretaceous (Cenomanian)	Sierra Madre Formation	Mexico	A close relative of Aipichthyoides and Aspesaipichthys, more distantly related to Aipichthys. The type species is Zoqueichthys carolinae.

References

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[KeatingetalJVP2012-1] Joseph N. Keating; Robert S. Sansom; Mark A. Purnell (2012). "A new osteostracan fauna from the Devonian of the Welsh Borderlands and observations on the taxonomy and growth of Osteostraci". Journal of Vertebrate Paleontology. 32 (5): 1002–1017. doi:10.1080/02724634.2012.693555.

[2] Zhu Min; Liu Yu-Hai; Jia Lian-Tao; Gai Zhi-Kun (2012). "A new genus of eugaleaspidiforms (Agnatha: Galeaspida) from the Ludlow, Silurian of Qujing, Yunnan, Southwestern China" (PDF). Vertebrata PalAsiatica. 50 (1): 1–7. doi:10.19615/j.cnki.1000-3118.2012.01.001.

[3] Lankester E. R. (1868). "Monograph of fishes of the Old Red Sandstone of Britain. Part I. The Cephalaspidae". The Palaeontographical Society,London.: 62pp.

[4] Henning Blom (2012). "New birkeniid anaspid from the Lower Devonian of Scotland and its phylogenetic implications". Palaeontology. 55 (3): 641–652. doi:10.1111/j.1475-4983.2012.01142.x.

[5] Bradley R. Scott; Mark V. H. Wilson (2012). "A new species of Waengsjoeaspis (Cephalaspidomorpha, Osteostraci) from the Early Devonian of northwestern Canada, with a redescription of W. nahanniensis and implications for growth, variation, morphology, and phylogeny". Journal of Vertebrate Paleontology. 32 (6): 1235–1253. doi:10.1080/02724634.2012.694514.

[6] Carole J. Burrow; Kate Trinajstic; John Long (2012). "First acanthodian from the Upper Devonian (Frasnian) Gogo Formation, Western Australia". Historical Biology. 24 (4): 349–357. doi:10.1080/08912963.2012.660150.

[7] Héctor Botella; Carlos Martínez-Pérez; Rodrigo Soler-Gijón (2012). "Machaeracanthus goujeti n. sp. (Acanthodii) from the Lower Devonian of Spain and northwest France, with special reference to spine histology" (PDF). Geodiversitas. 34 (4): 761–783. doi:10.5252/g2012n4a3. Archived from the original (PDF) on 2013-12-14. Retrieved 2013-12-14.

[8] Victor Voichyshyn; Hubert Szaniawski (2012). "Acanthodian jaw bones from Lower Devonian marine deposits of Podolia, Ukraine". Acta Palaeontologica Polonica. 57 (4): 879–896. doi:10.4202/app.2011.0079.

[MalyshkinaPJ-9] T. P. Malyshkina (2012). "New sharks of the genus Abdounia (Carcharhiniformes: Carcharhinidae) from the Upper Eocene of the Trans-Ural Region". Paleontological Journal. 46 (4): 392–399. doi:10.1134/S0031030112040053.

[GuinotetalJSP-10] Guillaume Guinot; Henri Cappetta; Charlie J. Underwood & David J. Ward (2012). "Batoids (Elasmobranchii: Batomorphii) from the British and French Late Cretaceous". Journal of Systematic Palaeontology. 10 (3): 445–474. doi:10.1080/14772019.2011.588255.

[Adnetetal-11] Sylvain Adnet; Henri Cappetta; Guillaume Guinot & Giuseppe Notarbartolo di Sciara (2012). "Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference". Zoological Journal of the Linnean Society. 166 (1): 132–159. doi:10.1111/j.1096-3642.2012.00844.x.

[12] David M. Martill & Nizar Ibrahim (2012). "Aberrant rostral teeth of the sawfish Onchopristis numidus from the Kem Kem beds (?early Late Cretaceous) of Morocco and a reappraisal of Onchopristis in New Zealand". Journal of African Earth Sciences. 64: 71–76. doi:10.1016/j.jafrearsci.2011.11.009.

[13] Gerard R. Case; Todd D. Cook; Mark V.H. Wilson & Paul D. Borodin (2012). "A new species of the sclerorhynchid sawfish Borodinopristis from the Campanian (Upper Cretaceous) of North Carolina, USA". Historical Biology: An International Journal of Paleobiology. 24 (6): 592–597. doi:10.1080/08912963.2012.663367.

[14] Dana J. Ehret; Bruce J. MacFadden; Douglas S. Jones; Thomas J. Devries; David A. Foster & Rodolfo Salas-Gismondi (2012). "Origin of the white shark Carcharodon (Lamniformes: Lamnidae) based on recalibration of the Upper Neogene Pisco Formation of Peru". Palaeontology. 55 (6): 1139–1153. doi:10.1111/j.1475-4983.2012.01201.x.

[15] Claire Behan; Gordon Walken & Gilles Cuny (2012). "A Carboniferous chondrichthyan assemblage from residues within a Triassic karst system at Cromhall quarry, Gloucestershire, England". Palaeontology. 55 (6): 1245–1263. doi:10.1111/j.1475-4983.2012.01193.x.

[AndreevandCuny-16] Plamen S. Andreev; Gilles Cuny (2012). "New Triassic stem selachimorphs (Chondrichthyes, Elasmobranchii) and their bearing on the evolution of dental enameloid in Neoselachii". Journal of Vertebrate Paleontology. 32 (2): 255–266. doi:10.1080/02724634.2012.644646.

[AMS20121elasmobranchs-17] Baomin Zhang; Xiaohong Chen; Long Cheng & Miao Zhang (2012). "Middle Triassic (Ladinian) elasmobranch scales from the southwestern Guizhou, China". Acta Micropalaeontologica Sinica. 29 (1): 52–61.

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[19] Henri Cappetta; Sylvain Adnet; Driss Akkrim; Mohamed Amalik (2014). "New Squalicorax species (Neoselachii: Lamniformes) from the Lower Maastrichtian of Ganntour phosphate deposit, Morocco". Palæovertebrata. 38 (2–e3): 1–13. doi:10.18563/pv.38.2.e3.

[IvanovetalJSP-20] Alexander Ivanov; Merlynd Nestell; Galina Nestell (2012). "New jalodontid chondrichthyans from the Middle Permian of West Texas, USA". Historical Biology. 24 (4): 359–368. doi:10.1080/08912963.2012.662229.

[Hodnettetal-21] John-Paul M. Hodnett; David K. Elliott; Tom J. Olson; James H. Wittke (2012). "Ctenacanthiform sharks from the Permian Kaibab Formation, northern Arizona". Historical Biology. 24 (4): 381–395. doi:10.1080/08912963.2012.683193.

[22] Todd D. Cook; Mark V. H. Wilson; Alison M. Murray; A. Guy Plint; Michael G. Newbrey; Michael J. Everhart (2013). "A high latitude euselachian assemblage from the early Turonian of Alberta, Canada". Journal of Systematic Palaeontology. 11 (5): 555–587. doi:10.1080/14772019.2012.707990.

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[25] T. Bor; T. Reinecke; S. Verschueren (2012). "Miocene Chondrichthyes from Winterswijk – Miste, the Netherlands". Palaeontos. 21: 1–136.

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[27] Victor Eduardo Pauliv; Eliseu Vieira Dias & Fernando Antonio Sedor (2012). "A new species of sphenacanthid (Chondrichthyes, Elasmobranchii) from the Rio do Rasto Formation (Paraná Basin), southern Brazil". Revista Brasileira de Paleontologia. 15 (3): 243–250. doi:10.4072/rbp.2012.3.01.

[28] Ivan J. Sansom; Neil S. Davies; Michael I. Coates; Robert S. Nicoll; Alex Ritchie (2012). "Chondrichthyan-like scales from the Middle Ordovician of Australia". Palaeontology. 55 (2): 243–247. doi:10.1111/j.1475-4983.2012.01127.x.

[29] Michael D. Gottfried; R. Ewan Fordyce; Seabourne Rust (2012). "A new billfish (Perciformes, Xiphioidei) from the late Oligocene of New Zealand". Journal of Vertebrate Paleontology. 32 (1): 27–34. doi:10.1080/02724634.2012.634471.

[PI12188-30] Werner Schwarzhans (2012). "Fish otoliths from the Paleocene of Bavaria (Kressenberg) and Austria (Kroisbach and Oiching-Graben)". Palaeo Ichthyologica. 12: 1–88.

[31] Yoshitaka Yabumoto; Yoshinori Hikida; Takanobu Nishino (2012). "Apsopelix miyazakii, a New Species of Crossognathid Fish (Teleostei) from the Upper Cretaceous of Hokkaido, Japan". Paleontological Research. 16 (1): 37–46. doi:10.2517/1342-8144-16.1.037.

[32] Jesús Alvarado-Ortega & Héctor Gerardo Porras-Múzquiz (2012). "The first American record of Aspidopleurus (Teleostei, Aulopiformes), from La Mula Quarry (Turonian), Coahuila State, Mexico". Revista Brasileira de Paleontologia. 15 (3): 251–263. doi:10.4072/rbp.2012.3.02.

[33] Orangel Aguilera; Alexander P. Marceniuk (2012). "Aspistor verumquadriscutis, a new fossil species of sea catfishes (Siluriformes; Ariidae) from the upper Miocene of Venezuela". Swiss Journal of Palaeontology. 131 (2): 265–274. doi:10.1007/s13358-012-0043-x.

[Bawitius-34] Grandstaff, B. S; Smith, J. B.; Lamanna M. C.; Lacovara, K. J.; Abdel-Ghani M. S. (2012). "Bawitius, gen. nov., a giant polypterid (Osteichthyes, Actinopterygii) from the Upper Cretaceous Bahariya Formation of Egypt". Journal of Vertebrate Paleontology. 32 (1): 17–26. doi:10.1080/02724634.2012.626823.

[35] Louis Taverne; Luigi Capasso (2012). "Les poissons crétacés de Nardò. 35°. Compléments à l'étude des halécostomes Belonostomus (Aspidorhynchiformes) et Pseudopycnodus (Pycnodontiformes)" (PDF). Bollettino del Museo Civico di Storia Naturale di Verona. 36: 25–44.

[36] Hans-Peter Schultze; John W. Reed (2012). "A tristichopterid sarcopterygian fish from the upper Middle Devonian of Nevada". Historical Biology: An International Journal of Paleobiology. 24 (4): 425–440. doi:10.1080/08912963.2012.673599.

[SJG501-37] Matthew Parfitt; Zerina Johanson; Sam Giles; Matt Friedman (2014). "A large, anatomically primitive tristichopterid (Sarcopterygii: Tetrapodomorpha) from the Late Devonian (Frasnian) Alves Beds, Upper Old Red Sandstone, Moray, Scotland". Scottish Journal of Geology. 50 (1): 79–85. doi:10.1144/sjg2013-013.

[38] Giorgio Carnevale; Theodore W. Pietsch (2012). "†Caruso, a new genus of anglerfishes from the Eocene of Monte Bolca, Italy, with a comparative osteology and phylogeny of the teleost family Lophiidae". Journal of Systematic Palaeontology. 10 (1): 47–72. doi:10.1080/14772019.2011.565083.

[Congrophichthus-39] Werner Schwarzhans; Gary L. Stringer (2020). "Fish otoliths from the late Maastrichtian Kemp Clay (Texas, USA) and the early Danian Clayton Formation (Arkansas, USA) and an assessment of extinction and survival of teleost lineages across the K-Pg boundary based on otoliths". Rivista Italiana di Paleontologia e Stratigrafia. 126 (2): 395–446. doi:10.13130/2039-4942/13425.

[40] A. M. Prokofiev (2012). "Oligocene eel from the Frauenweiler site (Germany)". Journal of Ichthyology. 52 (1): 11–18. doi:10.1134/S0032945212010109.

[perleididfishes-41] Bing-he Geng; Fan Jin; Fei-xiang Wu; Qiang Wang (2012). "New perleidid fishes from the Middle Triassic strata of Yunnan Province" (PDF). Geological Bulletin of China. 31 (6): 915–927. Archived from the original (PDF) on 2015-09-23. Retrieved 2013-12-14.

[42] Alison M. Murray; Izzet Hoşgör (2012). "An early Oligocene elopiform fish from a new locality in Eastern Anatolia, Turkey". Journal of Vertebrate Paleontology. 32 (2): 296–303. doi:10.1080/02724634.2012.648248.

[43] James C. Tyler & Alexandre F. Bannikov (2012). "A new species of puffer fish, Eotetraodon tavernei, from the Eocene of Monte Bolca, Italy (Tetraodontidae, Tetraodontiformes)". Studi e Ricerche Sui Giacimenti Terziari di Bolca, XIV - Miscellanea Paleontologica. 11: 51–58.

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[45] Alexandre F. Bannikov & James C. Tyler (2012). "Redescription of the Eocene of Monte Bolca, Italy, surgeon fish Pesciarichthys punctatus (Perciformes, Acanthuridae), and a new genus, Frigosorbinia, for P. baldwinae". Studi e Ricerche Sui Giacimenti Terziari di Bolca, XIV - Miscellanea Paleontologica. 11: 19–30.

[46] A.F. Bannikov; G. Carnevale (2012). "Frippia labroiformis n. gen. n. sp., a new perciform fish from the Eocene of Pesciara di Bolca, Italy". Bollettino della Società Paleontologica Italiana. 51 (3): 155–165.

[otoliths-47] Werner Schwarzhans; R. Paul Scofield; Alan J. D. Tennyson; Jennifer P. Worthy; Trevor H. Worthy (2012). "Fish remains, mostly otoliths, from the non-marine early Miocene of Otago, New Zealand". Acta Palaeontologica Polonica. 57 (2): 319–350. doi:10.4202/app.2010.0127.

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[51] E. M. Baykina (2012). "A new clupeid genus (Pisces, Clupeiformes, Clupeidae) from the Sarmatian of the Eastern Paratethys, Krasnodar Region". Paleontological Journal. 46 (3): 302–312. doi:10.1134/S0031030112030057.

[52] E. M. Baykina (2013). "A revision of Clupea doljeana Kramberger and Sarmatella vukotinovici (Kramberger) (Pisces, Clupeidae) from the Sarmatian of Croatia". Paleontological Journal. 47 (5): 523–532. doi:10.1134/S0031030113050043.

[53] A. F. Bannikov (2012). "The first record of the genus Isurichthys (Perciformes, Ariommatidae) in the Lower Oligocene of the Northern Caucasus". Paleontological Journal. 46 (2): 171–176. doi:10.1134/S0031030112020025.

[Kyphosichthys-54] Xu, G. -H.; F.- X., Wu. (2012). "A deep-bodied ginglymodian fish from the Middle Triassic of eastern Yunnan Province, China, and the phylogeny of lower neopterygians". Chinese Science Bulletin. 57 (1): 111–118. doi:10.1007/s11434-011-4719-1.

[Nardoelops-55] Louis Taverne; Luigi Capasso (2012). "Les poissons crétacés de Nardò. 34°. Nardoelops nybelini gen. et sp. nov. et Lebonichthys nardoensis sp. nov. (Teleostei, Elopomorpha)" (PDF). Bollettino del Museo Civico di Storia Naturale di Verona. 36: 3–24.

[Luoxiongichthys-56] Wen Wen; Qi-Yue Zhang; Chang-Yong Zhou; Jin-Yuan Huang; Zhong Qiang Chen; Michael J. Benton (2012). "A new genus of basal actinopterygian fish from the Anisian (Middle Triassic) of Luoping, Yunnan Province, Southwest China". Acta Palaeontologica Polonica. 57 (1): 149–160. doi:10.4202/app.2010.0089.

[57] Kerstin M. Schröder; Adriana López-Arbarello; Martin Ebert (2012). "Macrosemimimus, gen. nov. (Actinopterygii, Semionotiformes), from the Late Jurassic of Germany, England, and France". Journal of Vertebrate Paleontology. 32 (3): 512–529. doi:10.1080/02724634.2012.649626.

[58] Stefan Koerber (2012). "Mercediella nom. nov., a replacement name for Camposichthys Figueiredo & Silva Santos, 1991 (Pisces: Pycnodontiformes)" (PDF). Bonn Zoological Bulletin. 61 (1): 29–30.

[59] Cesar R. L. Amaral; Paulo M. Brito (2012). "A New Chanidae (Ostariophysii: Gonorynchiformes) from the Cretaceous of Brazil with Affinities to Laurasian Gonorynchiforms from Spain". PLoS ONE. 7 (5): e37247. doi:10.1371/journal.pone.0037247. PMC 3357423. PMID 22629376.

[60] Florian Witzmann; Rainer R. Schoch (2012). "A megalichthyid sarcopterygian fish from the Lower Permian (Autunian) of the Saar-Nahe Basin, Germany". Geobios. 45 (2): 241–248. doi:10.1016/j.geobios.2011.03.002.

[61] Fernanda E. Weiss; Luiz R. Malabarba; Maria Claudia Malabarba (2012). "Phylogenetic relationships of Paleotetra, a new characiform fish (Ostariophysi) with two new species from the Eocene-Oligocene of south-eastern Brazil". Journal of Systematic Palaeontology. 10 (1): 73–86. doi:10.1080/14772019.2011.565082.

[62] Eliseu Vieira Dias (2012). "A new deep-bodied fossil fish (Actinopterygii) from the Rio do Rasto Formation, Paraná Basin, Brazil" (PDF). Zootaxa. 3192: 1–23.

[63] F.J. de Figueiredo; V. Gallo; A.F.P. Delarmelina (2012). "A new protacanthopterygian fish from the Upper Cretaceous (Turonian) of the Pelotas Basin, southern Brazil". Cretaceous Research. 34: 116–123. doi:10.1016/j.cretres.2011.10.009.

[64] Andrew J. Wendruff; Mark V. H. Wilson (2012). "A fork-tailed coelacanth, Rebellatrix divaricerca, gen. et sp. nov. (Actinistia, Rebellatricidae, fam. nov.), from the Lower Triassic of Western Canada". Journal of Vertebrate Paleontology. 32 (3): 499–511. doi:10.1080/02724634.2012.657317.

[65] John Graf (2012). "A new Early Cretaceous coelacanth from Texas". Historical Biology: An International Journal of Paleobiology. 24 (4): 441–452. doi:10.1080/08912963.2012.696636.

[66] Alice M. Clement (2012). "A new species of long-snouted lungfish from the Late Devonian of Australia, and its functional and biogeographical implications". Palaeontology. 55 (1): 51–71. doi:10.1111/j.1475-4983.2011.01118.x.

[67] C. Lombardo; A. Tintori; D. Tona (2012). "A new species of Sangiorgioichthys (Actinopterygii, Semionotiformes) from the Kalkschieferzone of Monte San Giorgio (Middle Triassic; Meride, Canton Ticino, Switzerland)". Bollettino della Società Paleontologica Italiana. 51 (3): 203–212.

[68] Aydın Akbulut; Hüseyin Özdikmen (2012). "To the memory of Dr. Bobb Schaeffer, Schaefferius nom. nov., a new name for the preoccupied genus Marquesia Schaeffer, 1990 (Osteichtyes: Actinopterygii: Palaeonisciformes)". Munis Entomology & Zoology. 7 (2): 1272–1273.

[69] Jiang-Yong Zhang (2012). "A new species of Sinamia from western Liaoning, China" (PDF). Vertebrata PalAsiatica. 50 (4): 322–334. doi:10.19615/j.cnki.1000-3118.2012.04.002.

[70] Alexandre F. Bannikov & Roberto Zorzin (2012). "A new genus and species of deep-bodies squamipinne-like percoid fish (Perciformes) frrom (SIC) the Eocene of Bolca in Norhtern [sic] Italy". Studi e Ricerche Sui Giacimenti Terziari di Bolca, XIV - Miscellanea Paleontologica. 11: 31–42.

[71] Andrea Tintori; Zuo-Yu Sun; Cristina Lombardo; Da-Yong Jiang; Cheng Ji; Ryosuke Motani (2012). "A new "flying" fish from the upper Ladinian (Middle Triassic) of Wusha (Guizhou Province, southern China)" (PDF). Gortania. Geologia, Paleontologia, Paletnologia. 33: 39–50. Archived from the original (PDF) on 2013-12-19. Retrieved 2013-12-18.

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[Tinirau-73] Brian Swartz (2012). "A Marine Stem-Tetrapod from the Devonian of Western North America". PLoS ONE. 7 (3): e33683. doi:10.1371/journal.pone.0033683. PMC 3308997. PMID 22448265.

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